Eilenodontinae

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Eilenodontinae
Temporal range: Late Triassic–Turonian
Priosphenodon skeleton.png
Skeleton of Priosphenodon avelasi
Eilenodon tooth and jaw.jpg
Teeth and jaws of Eilenodon
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Rhynchocephalia
Suborder: Sphenodontia
Subfamily: Eilenodontinae
Rasmussen and Callison 1981
Genera

Eilenodontinae are an extinct clade of reptiles belonging to Sphenodontia. They are either considered a subgroup of Opisthodontia, [1] or Sphenodontidae. [2] They had deep jaws with broad, closely packed teeth with thick enamel and noticeable wear facets. They were likely herbivorous, and probably chewed with a proal (forward stroke) movement, with food shredded between the edges of opposing sharp-edged wear facets. Members of the group are known from South America, North America and Europe. [3] The earliest known member of the group, Sphenotitan , is known from the Late Triassic of South America. [2] while the youngest members are known from the Late Cretaceous of South America. [4] The group contains some of the largest known sphenodontians, with Priosphenodon suggested to be the largest known non aquatic sphenodontian, with an estimated body length of over 1 metre (3.3 ft). [5]

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<span class="mw-page-title-main">Therapsid</span> Clade of tetrapods including mammals

A therapsid is a member of the clade Therapsida, which is a major group of eupelycosaurian synapsids that includes mammals and their ancestors and relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including limbs that were oriented more underneath the body, as opposed to the sprawling posture of many reptiles and salamanders.

<span class="mw-page-title-main">Gondwanatheria</span> Extinct group of Mammaliaformes that lived during the Upper Cretaceous through the Miocene

Gondwanatheria is an extinct group of mammaliaforms that lived in parts of Gondwana, including Madagascar, India, South America, Africa and Antarctica during the Upper Cretaceous through the Paleogene. Until recently, they were known only from fragmentary remains. They are generally considered to be closely related to the multituberculates and likely the euharamiyidians, well known from the Northern Hemisphere, with which they form the clade Allotheria.

<span class="mw-page-title-main">Cynodont</span> Clade of therapsids

Cynodonts are eutheriodont therapsids that first appeared in the Late Permian, and extensively diversified after the Permian–Triassic extinction event. Mammals are cynodonts, as are their extinct ancestors and close relatives (Mammaliaformes), having evolved from advanced probainognathian cynodonts during the Late Triassic.

<span class="mw-page-title-main">Rhynchocephalia</span> Order of reptiles

Rhynchocephalia is an order of lizard-like reptiles that includes only one living species, the tuatara of New Zealand. Despite its current lack of diversity, during the Mesozoic rhynchocephalians were a speciose group with high morphological and ecological diversity. The oldest record of the group is dated to the Middle Triassic around 238 to 240 million years ago, and they had achieved a worldwide distribution by the Early Jurassic. Most rhynchocephalians belong to the group Sphenodontia ('wedge-teeth'). Their closest living relatives are lizards and snakes in the order Squamata, with the two orders being grouped together in the superorder Lepidosauria.

<span class="mw-page-title-main">Sphenodontidae</span> Family of reptiles

Sphenodontidae is a family within the reptile group Rhynchocephalia, comprising taxa most closely related to the living tuatara. Historically the taxa included within Sphenodontidae have varied greatly between analyses, and the group has lacked a formal definition. Cynosphenodon from the Jurassic of Mexico has consistently been recovered as a close relative of the tuatara in most analyses, with the clade containing the two and other very close relatives of the tuatara often called Sphenodontinae. The herbivorous Eilenodontinae, otherwise considered part of Opisthodontia, is considered to be part of this family in many recent studies as the sister group to Sphenodontinae. The earliest Sphenodontines are known from the Early Jurassic of North America, with other remains known from the Late Jurassic of Europe, the Late Cretaceous of South America and the Miocene-recent of New Zealand. Sphenodontines are characterised by a complete lower temporal bar caused by the fusion of a forward directed process (extension) of the quadrate/quadratojugal and the jugal, which was an adaptation for reducing stress in the skull during hard biting. Other synapomorphies of Sphenodontinae include the presence of nasal foramina, a posterodorsal process of the coronoid of the lower jaw, the present of caniniform successional teeth at the front of the jaws, the presence of flanges on the posterior parts of teeth at back of the lower jaw, and an expanded radial condyle on the humerus. Like modern tuatara, members of Sphenodontinae were likely generalists with a carnivorous/insectivorous diet.

<span class="mw-page-title-main">Dryolestida</span> Extinct order of mammals

Dryolestida is an extinct order of mammals, known from the Jurassic and Cretaceous. They are considered basal members of the clade Cladotheria, close to the ancestry of therian mammals. It is also believed that they developed a fully mammalian jaw and also had the three middle ear bones. Most members of the group, as with most Mesozoic mammals, are only known from fragmentary tooth and jaw remains.

<i>Clevosaurus</i> Extinct genus of reptiles

Clevosaurus is an extinct genus of rhynchocephalian reptile from the Late Triassic and the Early Jurassic periods. Species of Clevosaurus were widespread across Pangaea, and have been found on all continents except Australia and Antarctica. Five species of Clevosaurus have been found in ancient fissure fill deposits in south-west England and Wales, alongside other sphenodontians, early mammals and dinosaurs. In regards to its Pangaean distribution, C. hadroprodon is the oldest record of a sphenodontian from Gondwana, though its affinity to Clevosaurus has been questioned.

<i>Eilenodon</i> Extinct genus of reptiles

Eilenodon is an extinct genus of rhynchocephalian reptile from the Late Jurassic Morrison Formation of western North America, present in stratigraphic zone 4. The only known species of this genus is Eilenodon robustus. It was a member of a group of rhynchocephalians called the eilenodontines, which were large, herbivorous members of Rhynchocephalia, the order of reptiles which contains the modern tuatara (Sphenodon). The generic name "Eilenodon" is Greek for "packed teeth", in reference to its closely packed teeth. The specific name, "robustus", refers to the strong build of the jaws.

<span class="mw-page-title-main">Megaraptora</span> Extinct clade of dinosaurs

Megaraptora is a clade of carnivorous theropod dinosaurs. Its derived members, the Megaraptoridae are noted for their large hand claws and powerfully-built forelimbs, which are usually reduced in size in other large theropods.

<i>Sphenotitan</i> Extinct genus of reptiles

Sphenotitan is an extinct genus of rhynchocephalian reptile, known from the Late Triassic (Norian) Quebrada del Barro Formation of Argentina. It is the earliest known member of the herbivorous Elienodontinae, and the only one known from the Triassic. It was a large-sized sphenodontian, with an estimated skull length of over 10 centimetres (3.9 in). The skull is roughly triangular in shape, and had large upper temporal fenestrae. The region of the skull in front of the eye socket is short. The premaxillae forms beak, with a cutting edge similar to a chisel. The teeth of Sphenotitan, like other elienodontines, were large and wide, and designed for shredding vegetation, with blade-like palatal teeth on the roof of the mouth.

<i>Morrosaurus</i> Extinct genus of dinosaurs

Morrosaurus is an extinct genus of herbivorous elasmarian dinosaur that lived in the late Cretaceous in Antarctica. The only known species is the type Morrosaurus antarcticus.

Priosphenodon is an extinct, large herbiviorous eilenodontine rhynchocephalian known from the mid-Cretaceous (Albian-Turonian) of Argentina. It is one of the largest known sphenodontians.

<span class="mw-page-title-main">Opisthodontia (reptile)</span> Clade of reptiles

Opisthodontia is a proposed clade of sphenodontian reptiles, uniting Opisthias from the Late Jurassic-earliest Cretaceous of Europe and North America with the Eilenodontinae, a group of herbivorous sphenodontians known from the Late Triassic to Late Cretaceous.

<span class="mw-page-title-main">Clevosaurs</span> Family of reptiles

Clevosaurs are an extinct group of rhynchocephalian reptiles from the Triassic and Jurassic periods.

<span class="mw-page-title-main">Sapheosaur</span> Extinct group of reptiles

Sapheosaurs are an extinct group of rhynchocephalian reptiles from the Late Jurassic period. "Sapheosaurs" is an informal name for a group of rhynchocephalians closely related to the genus Sapheosaurus. It was first recognized as a group containing multiple genera by Hoffstetter in 1955. The group has sometimes been given a formal taxonomic name as the family Sapheosauridae, although in some analyses this group belongs to the family Sphenodontidae and thus cannot be assigned its own family. They were fairly advanced rhynchocephalians which may have had semiaquatic habits.

This list of fossil reptiles described in 2020 is a list of new taxa of fossil reptiles that were described during the year 2020, as well as other significant discoveries and events related to reptile paleontology that occurred in 2020.

Kawasphenodon is an extinct genus of sphenodontian reptile, known from the Late Cretaceous and Paleocene of Patagonia in South America. The type species, K. expectatus, was described in 2005 from jaw fragments found in late Campanian aged sediments in the Los Alamitos Formation, the jaw when complete was estimated to be 11 cm long, making it among the largest known sphenodontians. A second species, K. peligrensis, around 1/3 the size of the type species, was described in 2014 also from jaw fragments in early Paleocene (Danian) sediments of the Salamanca Formation, making it the youngest known definitive representative of Rhynchocephalia outside of New Zealand. In the original description, it was found to be a member of Sphenodontidae, in some other subsequent analyses it was found to be a member of Opisthodontia. A 2020 analysis of rhyncocephalian relationships found it to be outside Opisthodontia, and instead a member of the Sphenodontinae as the closest known relative of the tuatara, with an estimated divergence between the two genera in the Early Cretaceous. Other subsequent studies have endorsed its placement as a member of Sphenodontidae. Like most other rhynchocephalians, the teeth are acrodont, with a deep dentary, and it probably had an omnivorous habit.

This list of fossil reptiles described in 2024 is a list of new taxa of fossil reptiles that were described during the year 2024, as well as other significant discoveries and events related to reptile paleontology that occurred in 2024.

References

  1. Romo de Vivar, Paulo R.; Martinelli, Agustín G.; Schmaltz Hsiou, Annie; Soares, Marina Bento (2020-07-02). "A New Rhynchocephalian from the Late Triassic of Southern Brazil Enhances Eusphenodontian Diversity" . Journal of Systematic Palaeontology. 18 (13): 1103–1126. doi:10.1080/14772019.2020.1732488. ISSN   1477-2019. S2CID   216226211.
  2. 1 2 Simões, Tiago R.; Kinney-Broderick, Grace; Pierce, Stephanie E. (2022-03-03). "An exceptionally preserved Sphenodon-like sphenodontian reveals deep time conservation of the tuatara skeleton and ontogeny". Communications Biology. 5 (1): 195. doi: 10.1038/s42003-022-03144-y . ISSN   2399-3642. PMC   8894340 . PMID   35241764.
  3. Jones, Marc E. H.; Lucas, Peter W.; Tucker, Abigail S.; Watson, Amy P.; Sertich, Joseph J. W.; Foster, John R.; Williams, Ruth; Garbe, Ulf; Bevitt, Joseph J.; Salvemini, Floriana (June 2018). "Neutron scanning reveals unexpected complexity in the enamel thickness of an herbivorous Jurassic reptile". Journal of the Royal Society Interface. 15 (143): 20180039. doi:10.1098/rsif.2018.0039. ISSN   1742-5689. PMC   6030635 . PMID   29899156.
  4. Gentil, Adriel R.; Agnolin, Federico L.; Garcia Marsà, Jordi A.; Motta, Matias J.; Novas, Fernando E. (June 2019). "Bridging the gap: Sphenodont remains from the Turonian (Upper Cretaceous) of Patagonia. Palaeobiological inferences". Cretaceous Research. 98: 72–83. doi:10.1016/j.cretres.2019.01.016. S2CID   135429146.
  5. Apesteguía, Sebastián; Novas, Fernando E. (9 October 2003). "Large Cretaceous sphenodontian from Patagonia provides insight into lepidosaur evolution in Gondwana". Nature. 425 (6958): 609–612. Bibcode:2003Natur.425..609A. doi:10.1038/nature01995. PMID   14534584. S2CID   4425130.