Erbenochile

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Erbenochile
Temporal range: Emsian to Eifelian
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Erbenochile erbeni.jpg
E.erbeni
Scientific classification
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Genus:
Erbenochile
Type species
Odontochile (Erbenochile) erbeni
Alberti, 1981
Species
  • E. erbeniAlberti, 1981 (type), = Odontochile (Erbenochile) erbeni
  • E. issoumourensisChatterton & Gibb, 2010

Erbenochile is a genus of spinose phacopid trilobite, of the family Acastidae, found in Lower to Middle Devonian age rocks from Algeria and Morocco. [1] [2] Originally described from an isolated pygidium (the posterior body part or shield), [1] the first complete articulated specimen of E. erbeni revealed the presence of extraordinarily tall eyes:

Contents

"Straight-sided towers of lenses... with [up to] 18 lenses in a vertical file"

Fortey & Chatterton (2003) [3]

Number of lenses has been estimated at 560 [3] or 450 [4] incomplete preservation accounting for the uncertainty. A lens count of 18 lenses per file is unusually high (twice that of closely related genus) and accounts for the height of the eye, as opposed to a noticeable increase in the size of the individual lenses. [4] A recently found species (E. issoumourensis) has smaller eyes, with fewer files (33–35) and fewer lenses per file (13–14 max) than E. erbeni. [4] E. erbeni eyes allow full 360 degree coverage in the horizontal plane and were high enough to allow the trilobite to see backwards over its thorax. The presence of eye shades, blocking glare from over head, validates the suggestion that (at least some) trilobites were diurnal and not nocturnal,

"a detail of life habit which had previously been equivocal: Eyeshades are no use in the dark."

Fortey & Chatterton (2003) [3]

Distribution

Description

Erbenochile issoumourensis, rear view, collected near Foum Zgid, Morocco, late Emsian, 45mm body length Erbenochile issoumourensis rearview 1 CRF.jpg
Erbenochile issoumourensis, rear view, collected near Foum Zgid, Morocco, late Emsian, 45mm body length

Erbenochile is a trilobite with very large eyes, with 33–36 vertical rows of 13–19 lenses each. The lower edge of eye slightly overhangs the adjacent part of the free cheek. The most backward part of the glabella (or occipital ring), the rings of the thorax, and rings on the pygidium each have a prominent median spine. The thorax consists of 11 segments. The frontal band of each of the lateral parts of the thorax segments (in the so-called pleural region) are not adorned. The posterior bands have a row of distinct small spines. The large tailshield (or pygidium) has 11 pairs of spines along the edge, and one in the middle, and 10–11 pairs of well-defined segments. The frontal pleural bands of each segment on the pygidium is lower compared to the posterior pleural band, and lacks the small spines of the posterior pleural band. The posterior border of pygidium is longer medially than laterally. [5]

Taxonomy

Erbenochile was originally designated as a subgenus to Odontochile , the type of the subfamily Odontochilinae, which was later synonymised with the subfamily Dalmanitinae. Morzadec promoted Erbenochile to full genus status and moved it to Asteropyginae, by virtue of having pygidial spines, a character unknown in Odontochilinae. Erbenochile also differs from all other Asteropyginid genera in a number of characters. The much larger number of segments of the pygidium, as expressed in the number of axial rings, pleural rib pairs and pygidial spines (23 versus 11, and in Gourdonia 13 spines) is most conspicuous. Also, the pygidial border has a smooth connection with the pygidial spines, while in the Asteropyginae the connection with the pygidial lappets have relief. Furthermore, in Erbenochile the split of the lateral glabellar furrow and the second and third transglabellar furrows (confusingly called S2 and S1) are very deep, comparable the situation in Dalmanitinae. [6]

Key to the species

Erbenochile erbeni, eye, "sunshade" and spines Erbenochile eye.JPG
Erbenochile erbeni, eye, "sunshade" and spines
Erbenochile issoumourensis, lateral view of the head Erbenochile issoumourensis sideview lefteyesharp CRF.jpg
Erbenochile issoumourensis, lateral view of the head
1The eyebrow-like palpebral lobe extends laterally over the visual surface of the eyes, "shading the eye against the sun". The visual surface is a vertical half-cylinder, with 18 or 19 lenses per vertical row. Front of the headshield (or cephalon) sticking out to the front (or short spatulate) and ornated with a row of upward and very short spines. Largest spines on the axis more than half as high as the eyes. Upper Emsian. Algeria and Morocco.
E. erbeni
-The palpebral lobe does not extend laterally. The visual surface has an angle, with the diameter at the top of the eye smaller than at its base, with 13 or 14 lenses per vertical row. Front of the cephalon rounded triangular and without tubercles. Largest spines on the axis less than half as high as the eyes. Latest Emsian or early Eifelian. Morocco.
E. issoumourensis

Related Research Articles

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<i>Phacops</i> Genus of arthropods (fossil)

Phacops is a genus of trilobites in the order Phacopida, family Phacopidae, that lived in Europe, northwestern Africa, North and South America and China from the Late Ordovician until the very end of the Devonian, with a broader time range described from the Late Ordovician. It was a rounded animal, with a globose head and large eyes, and probably fed on detritus. Phacops is often found rolled up ("volvation"), a biological defense mechanism that is widespread among smaller trilobites but further perfected in this genus.

<i>Dalmanites</i> Extinct genus of trilobites

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<i>Huntoniatonia</i> Genus of trilobites

Huntoniatonia is genus of trilobites, an extinct group of marine arthropods of average to large size.

<i>Walliserops</i>

Walliserops is a genus of spinose phacopid trilobite, of the family Acastidae, found in Lower to Middle Devonian age rocks from the Anti-Atlas Mountains of Morocco. All species of Walliserops possess a three-pronged "trident" that protrudes from the glabella. Walliserops is most closely related to the genus Comura.

<i>Drotops</i> Extinct genus of trilobites

Drotops is a genus of trilobites from the order Phacopida, family Phacopidae that lived during the Eifelian of the Middle Devonian. It was described by Struve in 1990 under type species Drotops megalomanicus. Their fossils are found in present-day Morocco, specifically the Maïder Region located South West of Erfoud.

<i>Dikelocephalus</i> Genus of trilobites

Dikelocephalus is a genus of very large trilobites of up to 50 cm (20 in) long, that lived during the last 3 million years of the Cambrian (Sunwaptan). Their fossils are commonly found as disarticulated sclerites, in the upper Mississippi Valley and in Canada (Alberta). The exoskeleton is rounded anteriorly, with the thorax and sides of the tailshield slightly tapering to about ⅔× of the width across the base of the spines at the back of the headshield. At the side corners of the pygidium there may be triangular or hooked spines, pointing backwards, while between the spines the posterior margin is at a 30-75° angle with the lateral margin, gently convex or nearly straight. If pygidial spines are lacking, the margin is gradually rounded. The thorax has 12 segments.

<i>Tsunyidiscus</i> Genus of trilobites

Tsunyidiscus is a trilobite belonging to the Suborder Eodiscina. Tsunyidiscus appeared near the end of the Lower Cambrian, during the late Atdabanian stage of geologic time and some collections suggest it may have survived into the Botomian. The genus is very small, oculate and isopypous with a narrow dome-shaped glabella and a narrow bullet-shaped pygidial axis. Thorax consists of three segments. Tsunyidiscus is the only genus currently attributed to the family Tsunyidiscidae.

<i>Meteoraspis</i>

Meteoraspis is an extinct genus of ptychopariid trilobites of the family Tricrepicephalidae. The various species lived from 501 to 490 million years ago during the Dresbachian faunal stage of the late Cambrian Period. Fossils of Meteoraspis are characteristic of Late Cambrian strata in North America, though they are found in Late Cambrian strata elsewhere in the world, such as M. nevensis from Victoria Land, Antarctica.

Fenestraspis is an extinct genus of trilobite in the order Phacopida from the Upper Pragian and Lower Emsian. Fenestraspis is unusual because of the development of extensive fenestrae in the posterior part of the body and apparently of the thorax, the presence of upwardly directed spines on the cephalon, thorax and pygidium, and the exceptionally large and highly elevated eyes.

<i>Morocconites</i>

Morocconites malladoides is an average size trilobite, which lived during the Devonian period, in what is now southern Morocco. This species is assumed to be a close relative of Acastoides. The most conspicuous feature is the very long upcurved frontal medial spine, a bit like an avocet bill. It is the only known species in this genus.

<i>Odontochile</i> Genus of trilobites

Odontochile is a genus of trilobites in the order Phacopida, family Dalmanitidae.

Eodiscina

Eodiscina is trilobite suborder. The Eodiscina first developed near the end of the Lower Cambrian period and became extinct at the end of the Middle Cambrian. Species are tiny to small, and have a thorax of two or three segments. Eodiscina includes six families classified under one superfamily, Eodiscoidea.

<i>Pricyclopyge</i> Extinct genus of trilobites

Pricyclopyge is a genus of trilobites assigned to the family Cyclopygidae that occurs throughout the Ordovician. Pricyclopyge had an extratropical distribution, and there is evidence that it lived in darker parts of the water column. Pricyclopyge has huge eyes, an inverted pear-shaped glabella, six thorax segments, with on the 3rd two small discs. Pricyclopyge is known from what are today China, the Czech Republic, France, and the United Kingdom.

<i>Coltraneia</i>

Coltraneia is a genus of trilobite, that lived during the upper Emsian and lower Eifelian, and has been found in Algeria, France, Germany, Morocco and Spain.

<i>Minicryphaeus</i>

Minicryphaeus is a genus of trilobite, that lived during the Pragian in what is today Morocco (Anti-Atlas).

<i>Psychopyge</i>

Psychopyge is a genus of trilobite, that lived during the upper Emsian and has been found in Germany and Morocco. It is characterized by the swordlike extension from the front of the head.

<i>Placoparia</i> Genus of trilobites

Placoparia is a genus of trilobites of average size that lived during the late Lower to the early Upper Ordovician on the paleocontinents Gondwana, Avalonia and Laurentia, now the Czech Republic, France, Germany, Morocco, Portugal, Spain and Wales. Its headshield is semi-circular to rectangular with rounded frontal corners. It lacks eyes, but eye ridges are present. The fact that the facial sutures are opisthoparian is an exception in the otherwise proparian Cheirurina. The thorax has 11 or 12 segments, with the axis slightly wider than the ribs to its sides. The tips of the pleurae are free, which resembles an old-fashion central heating radiator. The axis in the small tailshield consists of four rings and a minute endpiece. The four pleurae end in spatulate spines that fit to corresponding indentations in the cephalon.

<i>Tricrepicephalus</i>

Tricrepicephalus is an extinct genus of ptychopariid trilobites of the family Tricrepicephalidae with species of average size. Its species lived from 501 to 490 million years ago during the Dresbachian faunal stage of the late Cambrian Period. Fossils of Tricrepicephalus are widespread in Late Cambrian deposits in North America, but is also known from one location in South-America. Tricrepicephalus has an inverted egg-shaped exoskeleton, with three characteristic pits in the fold that parallels the margin of the headshield just in front of the central raised area. The articulating middle part of the body has 12 segments and the tailshield carries two long, tubular, curved pygidial spines that are reminiscent of earwig's pincers that rise backwards from the plain of the body at approximately 30°.

<i>Viaphacops</i>

Viaphacops is a genus of trilobites in the order Phacopida, family Phacopidae, that lived during the Middle Devonian, and is known from North and South America, Asia.

References

  1. 1 2 Morzadec, P. (1995), "Erbenochile erbeni (Alberti), Trilobite de Dévonian inférieur d'Ougarta (Algerie).", Neues Jahrb. Geol. Paläontol. Monatsh., 10: 614–621
  2. Alberti, G.K.B. (1981), "Beziehungen zwischen "herzynischen" Trilobitenfauna NW-Marokko und Deutschland (Unter- und Mittel-Devon)", Natur und Museum, 110: 172–182
  3. 1 2 3 Fortey, R.; Chatterton, B. (2003), "A Devonian Trilobite with an Eyeshade", Science, 301: 1689, doi:10.1126/science.1088713, PMID   14500973
  4. 1 2 3 Chatterton, B.; Fortey, R.; Brett, K.; Gibbs, S.; McKellar, R. (2006), "Trilobites from the Lower to Middle Devonian Timrhanrhart Formation, Jbel Gara el Zquilma, southern Morocco.", Palaeontographica Canadiana, 25, ISBN   1-897095-15-5
  5. 1 2 Chatterton, B.D.E.; Gibb, S. (2010). "Latest Early and early Middle Devonian Trilobites from the Erbenochile bed, Jbel Issoumour, southeastern Morocco". Journal of Paleontology. 84 (6): 1188–1205. doi:10.1666/10-027.1 . Retrieved 31 December 2013.
  6. Bignon, Arnaud; Crônier, Catherine (2013). "The systematics and phylogeny of the Devonian subfamily Asteropyginae (Trilobita: Phacopida)". Journal of Systematic Palaeontology: 1–32. doi:10.1080/14772019.2013.804008. hdl: 11336/32076 .