Eurypterids, often informally called sea scorpions, are a group of extinct arthropods that form the order Eurypterida. The earliest known eurypterids date to the Darriwilian stage of the Ordovician period 467.3 million years ago. The group is likely to have appeared first either during the Early Ordovician or Late Cambrian period. With approximately 250 species, the Eurypterida is the most diverse Paleozoic chelicerate order. Following their appearance during the Ordovician, eurypterids became major components of marine faunas during the Silurian, from which the majority of eurypterid species have been described. The Silurian genus Eurypterus accounts for more than 90% of all known eurypterid specimens. Though the group continued to diversify during the subsequent Devonian period, the eurypterids were heavily affected by the Late Devonian extinction event. They declined in numbers and diversity until becoming extinct during the Permian–Triassic extinction event 251.9 million years ago.
Horseshoe crabs are marine and brackish water arthropods of the family Limulidae and the only living members of the order Xiphosura. Despite their name, they are not true crabs or crustaceans: they are chelicerates, most closely related to arachnids such as spiders, ticks, and scorpions.
Xiphosura is an order of arthropods related to arachnids. They are more commonly known as horseshoe crabs. They first appeared in the Hirnantian. Currently, there are only four living species. Xiphosura contains one suborder, Xiphosurida, and several stem-genera.
Willwerathia is a genus of Devonian arthropod. It is sometimes classified as synziphosurine, a paraphyletic group of horseshoe crab-like fossil chelicerate arthropods, while some studies compare its morphology to an artiopod. Willwerathia known only by one species, Willwerathia laticeps, discovered in deposits of the Devonian period from the Klerf Formation, in the Rhenish Slate Mountains of Germany.
Tarsopterella is a genus of prehistoric eurypterid classified within the family Hardieopteridae. It contains only one species, T. scotica from the Lower Devonian of Scotland.
Bembicosoma is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. Bembicosoma was regarded as part of the clade Planaterga. Fossils of the single and type species, B. pomphicus, have been discovered in deposits of the Silurian period in the Pentland Hills, Scotland. Bembicosoma had been tentatively assigned as an eurypterid before its synziphosurine affinities revealed.
Bunaia is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. Bunaia was tentatively placed as part of the clade Planaterga. The genus contains at least one species: Bunaia woodwardi from the Silurian period in Svalbard, Norway. Only a few morphological information of B. woodwardi had been confirmed, as the species known only from poorly preserved specimens compose of semicircular carapace, fragments of opisthosoma and disarticulated telson. The placement of "Bunaia" heintzi within this genus had been questioned and required further investigation.
Bunodes is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. Bunodes was regarded as part of the clade Planaterga. Fossils of the single and type species, B. lunula, have been discovered in deposits of the Silurian period in Ludlow, England. Bunodes is the type genus of the family Bunodidae, the other genera of the same family being Limuloides.
Legrandella is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. Legrandella was regarded as part of the clade Prosomapoda. Fossils of the single and type species, L. lombardii, have been discovered in deposits of the Devonian period in Cochabamba, Bolivia.
Limuloides is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. Limuloides was regarded as part of the clade Planaterga. Fossils of the genus have been discovered in deposits of the Silurian period in the United Kingdom and potentially in the United States. Limuloides is one of the two genera of the family Bunodidae, the other being the type genus Bunodes. Limuloides is characterized by a carapace with radiated ridges and serrated lateral regions, and an opisthosoma with rows of nodes. Limuloides was once though to have lateral compound eyes on its carapace, but later investigation did not find any evidence of it.
Pseudoniscus is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. Pseudoniscus was regarded as part of the clade Planaterga. Fossils of the genus have been discovered in deposits of the Silurian period in the United Kingdom, the United States and Estonia. Pseudoniscus is one of the two members of the family Pseudoniscidae, the other being Cyamocephalus.
Weinbergina is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. Fossils of the single and type species, W. opitzi, have been discovered in deposits of the Devonian period in the Hunsrück Slate, Germany.
Synziphosurina is a paraphyletic group of chelicerate arthropods previously thought to be basal horseshoe crabs (Xiphosura). It was later identified as a grade composed of various basal euchelicerates, eventually excluded form the monophyletic Xiphosura sensu stricto and only regarded as horseshoe crabs under a broader sense. Synziphosurines survived at least since early Ordovician to early Carboniferous in ages, with most species are known from the in-between Silurian strata.
The Pennine Coal Measures Group is a lithostratigraphical term referring to the coal-bearing succession of rock strata which occur in the United Kingdom within the Westphalian Stage of the Carboniferous Period. In formal use, the term replaces the Coal Measures Group as applied to the succession of coal-bearing strata within the Pennine Basin which includes all of the coalfields of northern England and the English Midlands. It includes the largely concealed Canonbie Coalfield of southern Scotland and the coalfields of northeast Wales and the minor Anglesey coalfield.
Austrolimulidae is an extinct family of horseshoe crabs belonging to the infraorder Limulina. Members of the family are known from the Permian to the beginning of the Jurassic, though one species has been reported from the end of the Cretaceous. Austrolimulids are known for amongst the most extreme morphologies among Xiphosurids, including large elongated genal spines. Unlike living Limulids, Austrolimulids were likely adapted for freshwater and brackish environments. They are considered to be the sister group to Limulidae, the group that contains all modern horseshoe crabs.
This timeline of eurypterid research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, and taxonomic revisions of eurypterids, a group of extinct aquatic arthropods closely related to modern arachnids and horseshoe crabs that lived during the Paleozoic Era.
Venustulus is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. Venustulus was regarded as part of the clade Prosomapoda. Fossils of the single and type species, V. waukeshaensis, have been discovered in deposits of the Silurian period in Wisconsin, in the United States. Venustulus is one of the few synziphosurine genera with fossil showing evidence of appendages, the other ones being Weinbergina, Anderella and Camanchia. Despite often being aligned close to horseshoe crabs, it has been found that Venustulus and its relatives form a group made up of various basal euchelicerate arthropods more distant to the xiphosurans.
Camanchia is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. Camanchia was regarded as part of the clade Prosomapoda. Fossils of the single and type species, C. grovensis, have been discovered in deposits of the Silurian period in Iowa, in the United States. Alongside Venustulus, Camanchia is one of the only Silurian synziphosurine with fossil showing evidence of appendages.
Anderella is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. Anderella was regarded as part of the clade Prosomapoda. Fossils of the single and type species, A. parva, have been discovered in deposits of the Carboniferous period in Montana, in the United States. Anderella is the first and so far the only Carboniferous synziphosurine being described, making it the youngest member of synziphosurines. Anderella is also one of the few synziphosurine genera with fossil showing evidence of appendages, but the details are obscure due to their poor preservation.
Maldybulakia is a genus of freshwater arthropod which lived during Late Silurian to Late Devonian. Maldybulakia is known from three species, M. angusi and M. malcolmi from Australia, M. mirabilis from Kazakhstan, and M. saierensis from China. Its classification is uncertain, it was originally described as a myriapod-like animal, and later considered related to the xiphosurans, or an artiopod.
