Geckos are small, mostly carnivorous lizards that have a wide distribution, found on every continent except Antarctica. Belonging to the infraorder Gekkota, geckos are found in warm climates throughout the world. They range from 1.6 to 60 centimetres.
Gekkonidae is the largest family of geckos, containing over 950 described species in 64 genera. The Gekkonidae contain many of the most widespread gecko species, including house geckos (Hemidactylus), the tokay gecko (Gekko), day geckos (Phelsuma), the mourning gecko (Lepidodactylus), and dtellas (Gehyra). Gekkonid geckos occur globally and are particularly diverse in tropical areas.
Dibamidae or blind skinks is a family of lizards characterized by their elongated cylindrical body and an apparent lack of limbs. Female dibamids are entirely limbless and the males retain small flap-like hind limbs, which they use to grip their partner during mating. They have a rigidly fused skull, lack pterygoid teeth and external ears. Their eyes are greatly reduced, and covered with a scale.
Xenosauridae is a family of anguimorph lizards whose only living representative is the genus Xenosaurus, which is native to Central America. Xenosauridae also includes the extinct genera Exostinus and Restes. Also known as knob-scaled lizards, they have rounded, bumpy scales and osteoderms. Most living species prefer humid, rocky habitats, although they are widespread within their native regions, with some inhabiting semi-arid scrub environments. They are carnivorous or insectivorous, and give birth to live young.
The Varanidae are a family of lizards in the superfamily Varanoidea and order Anguimorpha. The family, a group of carnivorous and frugivorous lizards, includes the living genus Varanus and a number of extinct genera more closely related to Varanus than to the earless monitor lizard (Lanthanotus). Varanus includes the Komodo dragon, crocodile monitor, savannah monitor, the goannas of Australia and Southeast Asia, and various other species with a similarly distinctive appearance. Their closest living relatives are the earless monitor lizard and chinese crocodile lizard. The oldest members of the family are known from the Late Cretaceous of Mongolia.
Scleroglossa is a group of lizards that includes geckos, autarchoglossans, and amphisbaenians. Scleroglossa is supported by phylogenetic analyses that use morphological features. According to most morphological analyses, Scleroglossa is the sister group of the clade Iguania, which includes iguanas, chameleons, agamids, and New World lizards. Together, Scleroglossa and Iguania make up the crown group Squamata, the smallest evolutionary grouping to include all living lizards and snakes.
Platynota is a polyphyletic group of anguimorph lizards and thus belongs to the order Squamata of the class Reptilia. Since it was named in 1839, it has included several groups, including monitor lizards, snakes, mosasaurs, and helodermatids. Its taxonomic use still varies, as it is sometimes considered equivalent to the group Varanoidea and other times viewed as a distinct group. It is phylogenetically defined as a clade containing Varanidae. It also includes many extinct species.
Dorsetisaurus is a genus of extinct lizard, known from the Late Jurassic of North America, and the Late Jurassic-earliest Cretaceous of Europe. The genus was first reported from the Early Cretaceous (Berriasian) Lulworth Formation of the Purbeck Group of Durlston Bay, in Dorset. It has also been reported from the Late Jurassic aged Alcobaça Formation of Portugal, the Aptian-Albian Dzunbain Formation of Mongolia, and the Morrison Formation of Western North America present in stratigraphic zones 2, 4, and 5. It is considered the oldest widely accepted member of Anguimorpha. based on the presence of 11 shared synapomorphies.
The Carphodactylidae, informally known as the southern padless geckos, are a family of geckos, lizards in the infraorder Gekkota. The family consists of 32 described species in 7 genera, all of which are endemic to Australia. They belong to the superfamily Pygopodoidea, an ancient group of east Gondwanan geckos now only found in Australasia. Despite their well-developed limbs, molecular phylogenies have demonstrated that Carphodactylidae is the sister group to Pygopodidae, a highly specialized family of legless lizards.
The Sphaerodactylidae are a family of geckos (Gekkota) distributed in North America, Central America, South America, and the Caribbean, as well as in Southern Europe, North Africa, the Middle East, and into Central Asia. The family contains 12 living genera and over 200 living species. The family name comes from the ball shape of their finger joints.
The Diplodactylidae are a family in the suborder Gekkota (geckos), with over 150 species in 25 genera. These geckos occur in Australia, New Zealand, and New Caledonia. Diplodactylids are the most ecologically diverse and widespread family of geckos in both Australia and New Caledonia, and are the only family of geckos found in New Zealand. Three diplodactylid genera have recently been split into multiple new genera.
Gobekko is an extinct genus of gecko or gecko-like lizard from the Late Cretaceous Djadokhta Formation of the Gobi Desert in Mongolia. Gobekko is either a basal member of Gekkota, the group that includes geckos and the legless pygopodid lizards, or a stem-gekkotan outside Gekkota but within the larger group Gekkonomorpha. It is the fourth oldest known member of Gekkonomorpha after Hoburogekko, a gecko from the Early Cretaceous of Mongolia, AMNH FR21444, an unnamed specimen also from the Early Cretaceous of Mongolia, and Cretaceogekko, a gecko preserved in amber from the Early Cretaceous of Burma.
Hoburogekko is an extinct genus of gecko that includes a single species, Hoburogekko suchanovi, from the Early Cretaceous Dzunbain Formation of Mongolia. It is known from two fossil specimens, one preserving the front part of the skull and the other preserving part of the lower jaw. Hoburogekko is one of four known Mesozoic geckos or gecko-like lizards, the others being Cretaceogekko from the Early Cretaceous of Burma, AMNH FR21444, an undescribed specimen from a slightly older deposit in Mongolia, and Gobekko from the Late Cretaceous of Mongolia. Hoburogekko is the third oldest known gecko behind AMNH FR21444 and Cretaceogekko.
Shinisauria is a clade or evolutionary grouping of anguimorph lizards that includes the living Chinese crocodile lizard Shinisaurus and several of its closest extinct relatives. Shinisauria was named in 2008 by Jack Lee Conrad as a stem-based taxon to include all anguimorphs more closely related to Shinisaurus than to Anguis fragilis, Heloderma suspectum or Varanus varius. Several recent phylogenetic analyses of lizard evolutionary relationships place Shinisauria in a basal position within the clade Platynota, which also includes monitor lizards, helodermatids, and the extinct mosasaurs. Shinisaurians were once thought to be closely related to the genus Xenosaurus, but they are now considered distant relatives within Anguimorpha. The fossil record of shinisaurians extends back to the Early Cretaceous with Dalinghosaurus, which is from the Aptian aged Yixian Formation of China. Two other extinct shinisaurians are currently known: Bahndwivici from the Eocene of Wyoming and Merkurosaurus from the Late Oligocene of Germany and the Early Miocene of the Czech Republic. An indeterminate shinisaurian is known from an isolated tail found in the Eocene aged Messel pit in Germany.
Myrmecodaptria is an extinct genus of scleroglossan lizard from the Late Cretaceous Djadokhta Formation in Ukhaa Tolgod, Mongolia. The type and only species, Myrmecodaptria microphagosa, was named in 2000 by paleontologists Gao Keqin and Mark Norell. Myrmecodaptria is known from a single holotype skull and lower jaws. It is distinguished from all other lizards by its extremely elongated skull. The eyes are placed close to the snout, which is short and rounded. The top of the skull is covered in bony knobs called osteoderms. The parietal bone at the back of the skull is elongated and about as long as the frontal bones, which are the usually the longest bones along the top of the skull in lizards. The squamosal bone at the back of the skull reaches forward to connect with the jugal bone behind the eye, forming a thin arch between the temporal fenestrae. Myrmecodaptria also has fewer and more widely spaced teeth in its jaws than do most other lizards.
Chamaeleontiformes is a hypothesized clade of iguanian lizards defined as all taxa sharing a more recent common ancestor with Chamaeleo chamaeleon than with Hoplocercus spinosus, Polychrus marmoratus, or Iguana iguana. It was named by paleontologist Jack Conrad in 2008 to describe a clade recovered in his phylogenetic analysis that included the extinct genus Isodontosaurus, the extinct family Priscagamidae, and the living clade Acrodonta, which includes agamids and chameleons. It is a stem-based taxon and one of two major clades within Iguania, the other being Pleurodonta. Below is a cladogram from Daza et al. (2012) showing this phylogeny:
Norellius is an extinct genus of scleroglossan lizard from the Early Cretaceous Öösh Formation of Mongolia. It is known from a well-preserved skull that was collected by an American Museum of Natural History expedition to Mongolia in 1923 and cataloged as AMNH FR 21444. After its initial cataloging, the specimen was not mentioned again in the scientific literature until 2004, when it was recognized as belonging to a potential early relative of modern groups of squamates such as gekkotans, amphisbaenians, dibamids, and snakes. AMNH FR 21444 was more fully described in a 2006 study that used high-resolution computed tomography to examine the skull and its braincase, and was described as a new genus and species, Norellius nyctisaurops, in 2015. The genus name honors paleontologist Mark Norell. The 2006 study incorporated the specimen into a phylogenetic analysis and found it to be a basal member of an evolutionary grouping called Gekkonomorpha, a stem-based taxon that includes living geckos and legless lizards (pygopodids) and all taxa more closely related to them than to any other living lizard. Norellius lies outside the node-based taxon Gekkota, a more strictly-defined subgroup of Gekkonomorpha that includes geckos, pygopodids, and all descendants of their most recent common ancestor. Therefore, while Norellius is more closely related to geckos and pygopodids than it is to any other living group of lizards, it branched off before the most recent common ancestor of these two groups. Below is a cladogram showing the position of Norellius according to this phylogeny:
Eichstaettisaurus is a genus of lizards from the Late Jurassic and Early Cretaceous of Germany, Spain, and Italy. With a flattened head, forward-oriented and partially symmetrical feet, and tall claws, Eichstaettisaurus bore many adaptations to a climbing lifestyle approaching those of geckoes. The type species, E. schroederi, is among the oldest and most complete members of the Squamata, being known by one specimen originating from the Tithonian-aged Solnhofen Limestone of Germany. A second species, E. gouldi, was described from another skeleton found in the Matese Mountains of Italy. Despite being very similar to E. schroederi, it lived much later, during the Albian stage. Fossils of both species show exceptional preservation due to deposition in low-oxygen marine environments.
Pygopodoidea is a gecko superfamily and the only taxon in the gekkotan subclade Pygopodomorpha. The clade includes three Australasian families: Diplodactylidae, Carphodactylidae, and Pygopodidae. Traditional gekkotan systematics had considered Diplodactylidae and Carphodactylidae as subfamilies of the family Gekkonidae, but recent molecular work have placed Pygopodidae within Gekkonidae making it paraphyletic. These analyses have shown support of Pygopodidae and Carphodactylidae being sister taxa, with Diplodactylidae occupying a basal position in Pygopodoidea.
Ardeosauridae is an extinct family of lizards known from the Late Jurassic of Germany and North America and Early Cretaceous of Mongolia, with other potential species elsewhere from Europe and Asia over the same time period.
