The Cyatheaceae are a family of ferns, the scaly tree ferns, one of eight families in the order Cyatheales in the Pteridophyte Phylogeny Group classification of 2016. Alternatively, the family may defined much more broadly as the only family in the Cyatheales, with the PPG I family treated as the subfamily Cyatheoideae. The narrower circumscription is used in this article.
Alsophila camerooniana, synonym Cyathea camerooniana, is a species of tree fern native to Sierra Leone, Cameroon, northern Angola and western Uganda, where it grows in montane forest at an altitude of 900–1200 m. The trunk is erect and 2–3 m tall. Fronds are pinnate and 2–3 m long. The rachis ranges in colour from dark to pale and has some hairs on the underside. The stipe is dark and is covered with scales throughout. Where scales have fallen, small warts are present. The scales are glossy brown and have a thin, dull edge. Sori occur at the forks of veins and are covered by thin indusia, which range in shape from cup- to saucer-like.
Alsophila doctersii, synonym Cyathea doctersii, is a species of tree fern endemic to Sumatra, where it grows in forest at an altitude of approximately 150 m. The trunk is erect and 2–3 m tall. Fronds are bi- or tripinnate and 2–3 m long. The stipe is covered with scattered flat, brown scales and some hairs. Sori occur near the fertile pinnule midvein and are covered by thin, brown indusia that are scale-like in appearance.
Alsophila fenicis, synonym Cyathea fenicis, is a species of tree fern native to the Philippines, Taiwan, and Orchid Island, where it grows in wet forest, forest margins and on hillsides. The trunk is erect, up to 1 m tall and about 6 cm in diameter. Fronds are tripinnate and 1.5–2 m long. Characteristically of this species, the lowest pinnae are usually reduced. The stipe is spiny and ranges in colour from brown to purple-dark brown. It bears two types of scales: long dark brown scales as well as minute brown ones. Occasionally the scales are pale. Sori are round and arranged in two rows, one on either side of the pinnule midvein. They are covered by very small indusia that resemble scales in appearance.
Alsophila geluensis, synonym Cyathea geluensis, is a species of tree fern native to central and eastern New Guinea as well as the Louisiade Archipelago, where it grows in mossy forest. In New Guinea itself, plants grow at an altitude of 1000–2000 m, however they are present at lower elevations of 700–900 m on associated islands. The trunk of this tree fern is erect and may be 5 m tall or more. Fronds are bi- or tripinnate and 1–2.5 m long. They are usually about ten live fronds present in the crown at once. The stipe may be warty and/or have short spines as well as many scattered scales towards the base. These scales are pale to dark and have dull, fragile edges. Sori occur near the fertile pinnule midvein and are protected by pale, thin indusia. A. geluensis is a variable taxon and further study is needed to determine whether it does not in fact represent a species complex.
Gymnosphaera gigantea, synonyms Alsophila gigantea and Cyathea gigantea, is a species of tree fern native to northeastern to southern India, Sri Lanka, Nepal to Myanmar, Thailand, Laos, Vietnam, the Malay Peninsula, as well as central Sumatra and western Java. It grows in moist open areas at an elevation of 600–1000 m. The trunk of this species is erect and may be as tall as 5 m or more. Fronds are bi- or tripinnate and usually 2–3 m long. The rachis is long, dark to black in colouration and rough in appearance after the fall of scales. These scales are dark brown, glossy and have a narrow paler margin and fragile edges. Sori are round and indusia absent.
Alsophila grevilleana, synonym Cyathea grevilleana, is a species of tree fern endemic to Jamaica, where it grows in moist gullies and on wooded hills in both calcareous and noncalcareous soils at an altitude of 200–1200 m. The trunk of this plant is erect, about 7 m tall, and 10–15 cm in diameter. It is characteristically clothed in old stipe bases, brown scales and blackish spines. Fronds are tripinnate, dark green in colour, and up to 4 m in length. The last pinnae are sometimes separated, forming a distinctive clump around the trunk apex. The rachis is yellow-brown and almost smooth. This species has a long, dark brown stipe with a few scattered spines. Sori are produced in four to six pairs along the pinnule midvein. They are protected by pale brown indusia that are cup-like in appearance.
Alsophila halconensis, synonym Cyathea halconensis, is a species of tree fern native to the islands of Luzon and Mindoro in the Philippines, where it grows in forest at an altitude of 1200–1700 m. The trunk of this plant is erect and 2–4 m tall. Fronds may be bi- or tripinnate and more than 1 m in length. Conical spines and scales cover the stipe. These scales are dark brown in colouration and have fragile edges. Sori occur near the fertile pinnule midvein. They are protected by thin, pale indusia.
Gymnosphaera denticulata, synonyms Alsophila acaulis, Alsophila denticulata and Cyathea hancockii, is a species of tree fern native to Japan, the Ryukyu Islands, Taiwan, southern China, Hong Kong, and Hainan. It grows in forest, on stream banks, and in forest margins at an elevation of about 600 m or higher. The specific epithet hancockii commemorates William Hancock (1847-1914), who collected numerous plants in Japan, China and Southeast Asia.
Alsophila henryi, synonym Cyathea henryi, is a species of tree fern native to India and Yunnan in China, where it grows submontane and montane forest at an altitude of 600–1200 m. The trunk of this plant is erect and may be 5–7 m tall or more. Fronds are bi- or tripinnate and usually 2–3 m in length. The rachis is smooth and dark, but occasionally has a few scattered scales. The stipe also has these scales. The scales are either small and pale with irregular fringed edges, or large and dark with a paler margin. Sori are borne on minor veins and lack indusia.
Alsophila heterochlamydea, synonym Cyathea heterochlamydea, is a little-known species of tree fern native to the islands of Luzon, Panay, Negros and Mindanao in the Philippines, where it grows in montane forest. The trunk of this plant is erect and usually up to 4 m tall or more. Fronds may be bi- or tripinnate and 1–2 m in length. The stipe is warty and/or bears short spines and scales. These scales are dark, glossy and have a narrow pale margin. Sori are borne near the fertile pinnule midvein and are protected by firm, brown indusia.
Alsophila hooglandii, synonym Cyathea hooglandii, is a species of tree fern native to the Western Highlands of New Guinea, where it grows in mossy forest at an altitude of about 3000 m. The trunk of this plant is erect, up to 3 m tall and approximately 10 cm in diameter. Fronds are tripinnate, 1–2 m in length and produced in two whorls of five to seven each. The stipe may be dull and warty or densely covered with scales. These scales range in colour from dark to medium brown and have a broad, paler margin as well as fragile edges. Perhaps the most distinguishing feature of this species are its sori, which are borne singly at the base of each tertiary pinnule on the midvein. They are protected by firm indusia that are saucer-like in appearance.
Gymnosphaera hornei is a species of tree fern in the Cyatheaceae family.
Alsophila khasyana, synonym Cyathea khasyana, is a species of tree fern. Its natural distribution extends from India to Myanmar, although it is absent from Sri Lanka. A. khasyana grows in forest at an altitude of 1400–1700 m. The trunk of this plant is erect and 5–7 m tall. Fronds may be bi- or tripinnate and 2–3 m in length. A. khasyana has a long, dark stipe that is covered in numerous scales. These scales are dark and have broad, pale, fringed edges. Sori are borne near the midvein of fertile pinnules and lack indusia.
Alsophila humilis, synonym Cyathea humilis, is a species of tree fern native to Kenya, as well as the Usambara and Uluguru Mountains in Tanzania, where it grows in wet forest at an altitude of 1100–2000 m. The trunk of this plant is erect and 2–3 m tall. Fronds are pinnate and 1–2 m in length. Dead fronds are often retained in the typical variety, forming an irregular skirt around the trunk. The rachis and stipe are light brown in colouration. Scales are present towards the base of the stipe. They are dark, glossy, and have narrow, fragile edges. Sori occur at the forks of veins and are protected by thin, reduced indusia.
Alsophila brausei, synonym Cyathea hunsteiniana, is a species of tree fern endemic to eastern New Guinea, where it grows in rain forest at an altitude of 1300–2000 m. This species has a slender, erect trunk up to 1 m tall and about 3 cm in diameter. Fronds are bipinnate and approximately 1 m in length. Scattered scales cover the stipe. They are dark brown in colouration and have a broad paler margin and fragile edges. Round sori are borne one or two per fertile pinnule segment. The sori are protected by deep, firm indusia that are cup-like in appearance.
Alsophila hymenodes, synonym Cyathea hymenodes, is a species of tree fern native to the Malay Peninsula, Sumatra, and possibly Java, where it grows in montane forest at an altitude of 900–2000 m. The trunk of this plant is erect and 2–4 m tall. Fronds may be bi- or tripinnate and 1–2 m in length. The stipe may have a pair of reduced pinnae towards the base and is covered with scattered scales. These scales are dark and have very fragile edges. Sori are borne near the fertile pinnule midvein and are protected by firm, brown indusia that are saucer-like in appearance.
Alsophila imbricata, synonym Cyathea imbricata, is a species of tree fern endemic to Western New Guinea, where it grows in open forest at an altitude of 3240 m. The trunk of this plant is erect and approximately 2 m tall. Fronds may be bi- or tripinnate and are usually less than 1 m in length. The stipe is dark, spiny, and covered with caducous scales. These scales are glossy brown in colouration and have a paler margin and fragile edges. Sori are borne in groups of one to four per pinnule lobe. They are protected by firm indusia.
Alsophila javanica, synonym Cyathea javanica, is a species of tree fern native to western Java and Sumatra, in Indonesia. It grows in rain forest and on riverbanks at an altitude of 250–1500 m.
Alsophila junghuhniana, synonym Cyathea junghuhniana, is a species of tree fern native to southern and central Sumatra and western Java, where it grows in forest at an altitude of 1000–2000 m. The trunk of this species is erect and may be 2 m tall or more. Fronds are tripinnate and up to about 3 m long. The stipes of this species are persistent and form a skirt around the trunk. They are brown, spiny, and covered with glossy, dark brown scales. Sori are borne near the midvein of fertile pinnules and are protected by thin indusia.
