Joculator

Joculator
Joculator variabilis
Scientific classification
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Subclass:	Caenogastropoda
Clade:	Hypsogastropoda
Suborder:	Ptenoglossa
Superfamily:	Triphoroidea
Family:	Cerithiopsidae
Genus:	Joculator Hedley, 1909
Type species
Cerithiopsis ridiculaWatson, 1886
Synonyms [1]
Cerithiopsis (Joculator)Hedley, 1909 Cyrbasia (Joculator)Hedley, 1909

Joculator is a genus of minute sea snails, marine gastropod molluscs in the family Cerithiopsidae. ^[1] This genus was described by Hedley in 1909.

Species

Species in the genus Joculator include:

• Joculator abbreviatus (Thiele, 1930)
• Joculator acuminatus Cecalupo & Perugia, 2012
• Joculator aeneus Cecalupo & Perugia, 2017
• Joculator albatus Cecalupo & Perugia, 2023
• Joculator albordina Laseron, 1956
• Joculator albulus (Thiele, 1930)
• Joculator alligatus Cecalupo & Perugia, 2012
• Joculator anconae Cecalupo & Perugia, 2017
• Joculator angustus Cecalupo & Perugia, 2017
• Joculator antonioi Cecalupo & Perugia, 2012
• Joculator aoreensis Cecalupo & Perugia, 2013
• Joculator arduinii Cecalupo & Perugia, 2012
• Joculator arenaceus Cecalupo & Perugia, 2013
• Joculator ariadnae Cecalupo & Perugia, 2018
• Joculator ater Cecalupo & Perugia, 2012
• Joculator aterrufus Cecalupo & Perugia, 2019
• Joculator autumnus Cecalupo & Perugia, 2016
• Joculator bangkaensis Cecalupo & Perugia, 2019
• Joculator baraggiai Cecalupo & Perugia, 2013
• Joculator barteri Cecalupo & Perugia, 2018
• Joculator barteri Cecalupo & Perugia, 2018
• Joculator bauquisi Cecalupo & Perugia, 2017
• Joculator berberiani Cecalupo & Perugia, 2018
• Joculator bicinctus Cecalupo & Perugia, 2012
• Joculator boguszae Cecalupo & Perugia, 2018
• Joculator bourgeoisae Cecalupo & Perugia, 2013
• Joculator boguszae Cecalupo & Perugia, 2018
• Joculator bouteti Cecalupo & Perugia, 2014 ^[2]
• Joculator brabantae Cecalupo & Perugia, 2013
• Joculator breviculus Cecalupo & Perugia, 2017
• Joculator brevis Cecalupo & Perugia, 2012
• Joculator brisseti Cecalupo & Perugia, 2021
• Joculator brucei (Melvill & Standen, 1912)
• Joculator brusonii Cecalupo & Perugia, 2013
• Joculator caliginosus Cecalupo & Perugia, 2012
• Joculator caloi Cecalupo & Perugia, 2013
• Joculator carpatinus Cecalupo & Perugia, 2012
• Joculator castaneus Cecalupo & Perugia, 2013
• Joculator cereus Cecalupo & Perugia, 2012
• Joculator christiaensi Jay & Drivas, 2002 ^[3]
• Joculator cinctus Cecalupo & Perugia, 2012
• Joculator citrinus Cecalupo & Perugia, 2013
• Joculator coffeus Cecalupo & Perugia, 2013
• Joculator conformis Cecalupo & Perugia, 2013
• Joculator continens Laseron, 1956
• Joculator corneliae Cecalupo & Perugia, 2018
• Joculator cossignanii Cecalupo & Perugia, 2012
• Joculator cristinae Cecalupo & Perugia, 2013
• Joculator czubai Cecalupo & Perugia, 2018
• Joculator dakoi Cecalupo & Perugia, 2018
• † Joculator decapitatus Grant-Mackie & Chapman-Smith, 1971
• Joculator desmieri Cecalupo & Perugia, 2018
• Joculator diversus Cecalupo & Perugia, 2019
• Joculator drivasi Cecalupo & Perugia, 2012
• Joculator dupouxae Cecalupo & Perugia, 2018
• Joculator emidioi Cecalupo & Perugia, 2014
• Joculator erianae Cecalupo & Perugia, 2017
• Joculator eudeli Jay & Drivas, 2002
• Joculator faninozi Cecalupo & Perugia, 2018
• Joculator fedosovi Cecalupo & Perugia, 2018
• Joculator ferrii Cecalupo & Perugia, 2013
• Joculator ferrugineus Cecalupo & Perugia, 2012
• Joculator filum Cecalupo & Perugia, 2017
• Joculator fischeri Jay & Drivas, 2002
• Joculator flindersi Cotton, 1951
• Joculator flavicans Cecalupo & Perugia, 2014
• Joculator frequens Cecalupo & Perugia, 2012
• Joculator fulvaster Cecalupo & Perugia, 2019
• Joculator furvus Cecalupo & Perugia, 2012
• Joculator fuscostriatus Cecalupo & Perugia, 2013
• Joculator fuscus Cecalupo & Perugia, 2012
• Joculator gagulae Cecalupo & Perugia, 2018
• Joculator ganui Cecalupo & Perugia, 2018
• Joculator garianii Cecalupo & Perugia, 2013
• Joculator gemmae Cecalupo & Perugia, 2012
• Joculator giovanolii Cecalupo & Perugia, 2013
• Joculator gorini Cecalupo & Perugia, 2014
• Joculator goubini Cecalupo & Perugia, 2017
• Joculator gracilis Laseron, 1951
• Joculator granatus Kay, 1979
• Joculator granulosus Cecalupo & Perugia, 2013
• Joculator grimalae Cecalupo & Perugia, 2021
• Joculator hadfieldae Cecalupo & Perugia, 2017
• Joculator hasegawai Cecalupo & Perugia, 2019
• Joculator hedleyi Laseron, 1951
• Joculator herosae Cecalupo & Perugia, 2012
• Joculator humilis Cecalupo & Perugia, 2012
• Joculator ianthinus Cecalupo & Perugia, 2013
• Joculator incisus Cecalupo & Perugia, 2012
• Joculator inflatus Cecalupo & Perugia, 2012
• Joculator introspectus Cotton, 1951
• Joculator iopuei Cecalupo & Perugia, 2017
• Joculator itiensis Cecalupo & Perugia, 2014
• Joculator jeffkinchi Cecalupo & Perugia, 2018
• Joculator johnkasui Cecalupo & Perugia, 2018
• Joculator juliuschani Cecalupo & Perugia, 2018
• Joculator kaipuae Cecalupo & Perugia, 2018
• Joculator kanoi Cecalupo & Perugia, 2013
• Joculator kenganziki Cecalupo & Perugia, 2018
• Joculator keratochroma Jay & Drivas, 2002
• Joculator keszlerae Cecalupo & Perugia, 2021
• Joculator koluae Cecalupo & Perugia, 2021
• Joculator kuboi Cecalupo & Perugia, 2019
• Joculator laguncula (Cecalupo & Perugia, 2012)
• Joculator laregnereensis Cecalupo & Perugia, 2017
• Joculator laseroni Jay & Drivas, 2002
• Joculator latrechei Cecalupo & Perugia, 2021
• Joculator lazzarii Cecalupo & Perugia, 2013
• Joculator legallae Cecalupo & Perugia, 2014
• Joculator legoffi Cecalupo & Perugia, 2017
• Joculator lazzarii Cecalupo & Perugia, 2013
• Joculator leguyaderi Cecalupo & Perugia, 2013
• Joculator lividus Cecalupo & Perugia, 2012
• Joculator lokaroensis Cecalupo & Perugia, 2014
• Joculator lozoueti Jay & Drivas, 2002
• Joculator luteolus Cecalupo & Perugia, 2012
• † Joculator maestratii Lozouet, Lesport & Renard, 2001
• Joculator magninensis Cecalupo & Perugia, 2017
• Joculator maloensis Cecalupo & Perugia, 2013
• Joculator manneensis Cecalupo & Perugia, 2018
• Joculator maranii Cecalupo & Perugia, 2014
• Joculator marinae Cecalupo & Perugia, 2014
• Joculator massimilianoi Cecalupo & Perugia, 2012
• Joculator mauberti Cecalupo & Perugia, 2018
• Joculator mbereensis Cecalupo & Perugia, 2017
• Joculator meanii Cecalupo & Perugia, 2014
• Joculator megacephala Jay & Drivas, 2002
• Joculator melania Laseron, 1956
• Joculator melanoraphis Jay & Drivas, 2002
• Joculator micalii Cecalupo & Perugia, 2012
• Joculator michoni Cecalupo & Perugia, 2018
• Joculator minimului Cecalupo & Perugia, 2018
• Joculator minimus Laseron, 1956
• Joculator minor Laseron, 1956
• Joculator minutissimus (Thiele, 1925)
• Joculator minutus Cecalupo & Perugia, 2012
• Joculator mitsuoi Cecalupo & Perugia, 2019
• Joculator modestus Cecalupo & Perugia, 2012
• Joculator morenae Cecalupo & Perugia, 2013
• Joculator morigii Cecalupo & Perugia, 2017
• Joculator murciai Cecalupo & Perugia, 2014
• Joculator mygaki Jay & Drivas, 2002
• Joculator myia Jay & Drivas, 2002
• Joculator nitidus Cecalupo & Perugia, 2012
• Joculator nucleus Cecalupo & Perugia, 2019
• Joculator obscurus Cecalupo & Perugia, 2012
• Joculator obsoletus Cecalupo & Perugia, 2012
• Joculator occultus Cecalupo & Perugia, 2012
• Joculator olivoideus Cecalupo & Perugia, 2018
• Joculator onnaensis Cecalupo & Perugia, 2019
• Joculator opulentus Cecalupo & Perugia, 2013
• Joculator ovatus Laseron, 1956
• Joculator pallidus Cecalupo & Perugia, 2012
• Joculator parvulus Cecalupo & Perugia, 2012
• Joculator pauxillus Cecalupo & Perugia, 2012
• Joculator perforatus Cecalupo & Perugia, 2013
• Joculator perlucidus Cecalupo & Perugia, 2012
• Joculator phtyr Jay & Drivas, 2002
• Joculator pierresamueli Cecalupo & Perugia, 2021
• Joculator pinguis Cecalupo & Perugia, 2012
• Joculator pizzinii Cecalupo & Perugia, 2013
• Joculator priorai Cecalupo & Perugia, 2012
• Joculator problematicus Albano & Steger, 2021
• Joculator prunus Cecalupo & Perugia, 2013
• Joculator psyllos Jay & Drivas, 2002
• Joculator pulvis (Issel, 1869)
• Joculator pumilus Cecalupo & Perugia, 2018
• Joculator pupiformis Cecalupo & Perugia, 2012
• Joculator pygmaeus Cecalupo & Perugia, 2012
• Joculator quaggiottoi Cecalupo & Perugia, 2012
• Joculator ralijanoai Cecalupo & Perugia, 2014
• Joculator ralphmanai Cecalupo & Perugia, 2018
• Joculator rambellii Cecalupo & Perugia, 2018
• Joculator recisus Cecalupo & Perugia, 2012
• Joculator ridiculus (Watson, 1886)
• Joculator rolani Cecalupo & Perugia, 2012
• Joculator rubus Cecalupo & Perugia, 2013
• Joculator sabrinae Cecalupo & Perugia, 2012
• Joculator saguili Cecalupo & Perugia, 2013
• Joculator salvati Jay & Drivas, 2002
• Joculator sarahpezetae Cecalupo & Perugia, 2018
• Joculator savellii Cecalupo & Perugia, 2021
• Joculator sbranai Cecalupo & Perugia, 2021
• Joculator schiaparellii Cecalupo & Perugia, 2017
• Joculator sekensis Cecalupo & Perugia, 2018
• Joculator semiperlucidus Cecalupo & Perugia, 2013
• Joculator simonsili Cecalupo & Perugia, 2018
• Joculator simulans Cecalupo & Perugia, 2012
• Joculator siwisikae Cecalupo & Perugia, 2018
• Joculator steykerae Cecalupo & Perugia, 2013
• Joculator stramineus Cecalupo & Perugia, 2013
• Joculator subconicus Cecalupo & Perugia, 2012
• Joculator subdolus Cecalupo & Perugia, 2012
• Joculator subglobosus Cecalupo & Perugia, 2013
• Joculator sublima Marshall, 1978
• Joculator subula Laseron, 1956
• Joculator succineus Cecalupo & Perugia, 2013
• Joculator sylviakinchae Cecalupo & Perugia, 2018
• Joculator tagliaferroi Cecalupo & Perugia, 2017
• Joculator testaceus Cecalupo & Perugia, 2013
• Joculator testii Cecalupo & Perugia, 2017
• Joculator theresae Cecalupo & Perugia, 2018
• Joculator thielei Jay & Drivas, 2002
• Joculator tieensis Cecalupo & Perugia, 2017
• Joculator tomacula Laseron, 1956
• Joculator tribulationis (Hedley, 1909)
• Joculator tsiriveloi Cecalupo & Perugia, 2014
• Joculator unicolor Cecalupo & Perugia, 2012
• Joculator uveanus (Melvill & Standen, 1896)
• Joculator vandelae Cecalupo & Perugia, 2014
• Joculator variabilis Cecalupo & Perugia, 2012
• Joculator varians Laseron, 1956
• Joculator vassardi Cecalupo & Perugia, 2014
• Joculator vazzanai Cecalupo & Perugia, 2018
• Joculator venustus Cecalupo & Perugia, 2013
• Joculator vignali Jay & Drivas, 2002
• Joculator violaceus Cecalupo & Perugia, 2012
• Joculator voncoseli Cecalupo & Perugia, 2012
• Joculator websterae Cecalupo & Perugia, 2014
• Joculator zahariasi Cecalupo & Perugia, 2021
• Joculator ziliolii Cecalupo & Perugia, 2012
• Joculator zucconi Cecalupo & Perugia, 2021

Taxa inquirenda

• Joculator aelomitres(Melvill & Standen, 1896)
• Joculator marileutes(Melvill & Standen, 1896)

Species brought into synonymy

• Joculator albocinctum(Melvill & Standen, 1896): synonym of Costulopsis albocincta (Melvill & Standen, 1896)
• Joculator albula (Thiele, 1930): synonym of Joculator albulus (Thiele, 1930)
• Joculator caelata Powell, 1930: synonym of Synthopsis caelata (Powell, 1930)
• Joculator granata Kay, 1979: synonym of Nanopsis granata (Kay, 1979)
• Joculator introspecta Cotton, 1951: synonym of Joculator introspectus Cotton, 1951
• Joculator iohannaeCecalupo & Perugia, 2012: synonym of Synthopsis iohannae (Cecalupo & Perugia, 2012)
• Joculator minima Laseron, 1956: synonym of Joculator minimus Laseron, 1956
• Joculator minutissima (Thiele, 1925): synonym of Joculator minutissimus (Thiele, 1925)
• Joculator myia Jay & Drivas, 2002: synonym of Nanopsis myia (Jay & Drivas, 2002)
• Joculator nanusLaseron, 1951: synonym of Potenatomus nanus (Laseron, 1951)
• Joculator ovata Laseron, 1956: synonym of Joculator ovatus Laseron, 1956
• Joculator ridicula: synonym of Joculator ridiculus (Watson, 1886)
• Joculator semipicta (Gould, 1861): synonym of Joculator semipictus (Gould, 1861)
• Joculator semipictus(Gould, 1861): synonym of Horologica semipicta (Gould, 1861)
• Joculator skolixJay & Drivas, 2002: synonym of Costulopsis skolix (Jay & Drivas, 2002) (original combination)
• Joculator turriger(Watson, 1886): synonym of Horologica turrigera (Watson, 1886)
• Joculator turrigera (Watson, 1886): synonym of Joculator turriger (Watson, 1886)
• Joculator uveanum (Melvill & Standen, 1896): synonym of Joculator uveanus (Melvill & Standen, 1896)

References

1. Rosenberg, G.; Bouchet, P. (2014). Joculator Hedley, 1909. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=531816 on 2014-12-07
2. "The Cerithiopsidae (Caenogastropoda: Triphoroidea) of South Madagascar (Indian Ocean)". Bollettino malacologico. 50 (2): 75–126. 2014. ISSN   0394-7149. Wikidata   Q135482828.
3. Maurice Jay; Jean Drivas (2002). "The Cerithiopsidae (Gastropoda) of Reunion Island (Indian Ocean)". Novapex. 3: 1–46. ISSN   1375-7474. Wikidata   Q135476558.

• Cecalupo A. & Perugia I., 2023. Analysis of Cerithiopsidae of the Jousseaume's collection. Xenophora Taxonomy 38: 17-24