Khodzhakul Formation

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Khodzhakul Formation
Stratigraphic range: Cenomanian
~93–89  Ma
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S
D
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Type Geological formation
Underlies Tamdytruba & Bissekty Formations
Overlies Bortesken Formation
Lithology
Primary Sandstone
Location
Coordinates 40°06′N65°48′E / 40.1°N 65.8°E / 40.1; 65.8
Approximate paleocoordinates 35°54′S65°30′E / 35.9°S 65.5°E / -35.9; 65.5
Region Karakalpakstan
Country Uruguay
Relief Map of Uzbekistan.png
Lightgreen pog.svg
Khodzhakul Formation (Uzbekistan)

The Khodzhakul Formation is a Cenomanian aged geologic formation in Uzbekistan. Dinosaur remains are among the fossils that have been recovered from the formation. [1] As well as those of Mammals. [2] It is part of the same stratigraphic succession as the overlying Bissekty Formation.

Contents

Fossil content

One of the largest mammals of the Mesozoic, Oxlestes , occurred in this formation, and may have predated on dinosaur species. [3]

Dinosaurs
GenusSpeciesLocationStratigraphic positionMaterialNotesImages

Asiaceratops [4]

A. salsopaludalis [4]

"Teeth, cranial fragments, phalanx." [5]

Gilmoreosaurus [6]

G. atavus [6]

Horezmavis [7]

H. eocretaceous [7]

"Tarsometatarsus." [8]

Kulceratops [4]

K. kulensis [4]

"Fragmentary material." [5]

Pectinodon [4]

P. sp. [4]

Therizinosauroidea

Indeterminate [9]

  • Upper

Ulughbegsaurus [10]

U. sp.

A right maxilla fragment

Mammals [2]
GenusSpeciesLocationStratigraphic positionMaterialNotesImages
Bobolestes B. zengeMaxilla and dentary fragmentsZalambdalestoidea
Eozhelestes E. mangitdentary fragments and teeth
Sheikhdzheilia S. rezvyiiMaxilla and dentary fragments?Zhelestidae
Zalambdalestidae IndeterminateRight petrosal
‘‘Zhelestidae’’IndeterminateLeft dentary and dentary fragmentLarge taxon
Color key
Taxon Reclassified taxonTaxon falsely reported as presentDubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

See also

References

  1. ↑ Weishampel, et al., 2004, "Dinosaur distribution." pp.517-607
  2. 1 2 Averianov, Alexander; Archibald, J. David (August 2005). "Mammals from the mid-Cretaceous Khodzhakul Formation, Kyzylkum Desert, Uzbekistan" (PDF). Cretaceous Research. 26 (4): 593–608. Bibcode:2005CrRes..26..593A. doi:10.1016/j.cretres.2005.03.007.
  3. ↑ Michael J. Benton,Mikhail A. Shishkin, David M. Unwin, The Age of Dinosaurs in Russia and Mongolia, Cambridge University Press, 04/12/2003 - 740 páginas
  4. 1 2 3 4 5 6 7 8 9 10 "40.3 Qoraqalpoghiston Respublikasi, Uzbekistan; 1. Upper Khodzhakul Formation," in Weishampel, et al. (2004). Page 594.
  5. 1 2 "Table 22.1," in Weishampel, et al. (2004). Page 480.
  6. 1 2 Listed as "? Gilmoreosaurus sp." in "40.3 Qoraqalpoghiston Respublikasi, Uzbekistan; 1. Upper Khodzhakul Formation," in Weishampel, et al. (2004). Page 594.
  7. 1 2 3 "40.3 Qoraqalpoghiston Respublikasi, Uzbekistan; 1. Lower Khodzhakul Formation," in Weishampel, et al. (2004). Page 563.
  8. ↑ "Table 11.1," in Weishampel, et al. (2004). Page 214.
  9. ↑ Sues, Hans-Dieter; Averianov, Alexander (2016). "Therizinosauroidea (Dinosauria: Theropoda) from the Upper Cretaceous of Uzbekistan". Cretaceous Research. 59: 155–178. Bibcode:2016CrRes..59..155S. doi: 10.1016/j.cretres.2015.11.003 .
  10. ↑ Averianov, A. O.; Kuzmin, I. T.; Skutschas, P. P.; Sues, H.-D. (2025). "First record of Carcharodontosauridae (Dinosauria, Theropoda) in the Upper Cretaceous Khodzhakul Formation of Uzbekistan". Journal of Paleontology: 1–13. doi:10.1017/jpa.2025.1.

Bibliography

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