| Myriopteris scabra | |
|---|---|
 | |
Scientific classification  | |
| Kingdom: | Plantae |
| Clade: | Tracheophytes |
| Division: | Polypodiophyta |
| Class: | Polypodiopsida |
| Order: | Polypodiales |
| Family: | Pteridaceae |
| Genus: | Myriopteris |
| Species: | M. scabra |
| Binomial name | |
| Myriopteris scabra (C.Chr.) Grusz & Windham | |
| Synonyms [1] | |
| |
Myriopteris scabra, commonly known as rough lipfern, is a species of cheilanthoid fern native to Mexico and the south-western United States (Texas). [2]
Myriopteris scabra grows from a short creeping rhizome, usually 4–7 mm in diameter with brown scales. The leaves (fronds) are clustered and may range greatly in size from 5–30 cm long. The leaf petiole is black to dark brown. The leaf blade is 1–4 cm wide, linear-oblong to lanceolate, and up to pinnate-pinnatifid to 2-pinnate. The rachis has scattered linear-lanceolate scales and dimorphic pubescence, abaxially sparsely hirsute, adaxially covered with tortuous appressed hairs. The ultimate leaflets are narrowly elliptic to elongate-deltate, not beadlike, and up to 3–5 mm long. The upper leaflet surface has a distinctive rough or spiky surface, which distinguishes this species from most other Myriopteris , and gives rise to the specific name scabra (Latin for rough or coarse). The leaflet edges are only folded under at their margins, barely concealing the sori (when present), which form a continuous bead around leaflet margins. [3]
Myriopteris scabra is native to northern Mexico, the Yucatan Peninsula, and to central and western Texas, extending north into Oklahoma. It grows on rocks and in rock crevices, mostly restricted to limestone, at elevations from 100 to 1400 meters. [3] Some close relatives of Myriopteris scabra are also calcareous soil/limestone specialists, including Myriopteris alabamensis , Myriopteris aemula , and Myriopteris microphylla . [4] [5] These species have ranges in the southern United States, Mexico, and Caribbean islands. Interestingly, another Myriopteris limestone specialist Myriopteris gracilis is not closely related to Myriopteris scabra. M. gracilis has a range that includes high mountains in the southwest United States and extends to much colder regions including Montana, Idaho, and British Columbia. [6]
According to plastid DNA sequence analysis, Myriopteris scabra is in the alabamensis clade of Myriopteris and its closest relative is Myriopteris fimbriata . [7]
Myriopteris covillei, formerly known as Cheilanthes covillei, is a species of cheilanthoid fern known by the common name Coville's lip fern. Coville's lip fern is native to the southwestern United States and Mexico.
Myriopteris parryi, formerly known as Cheilanthes parryi, is a species of lip fern known by the common name Parry's lip fern.
Myriopteris clevelandii, formerly known as Cheilanthes clevelandii, is a species of lip fern known by the common name Cleveland's lip fern. It is native to southern California and Baja California in Mexico. The leaf is divided into small, bead-like segments densely covered with scales beneath. In M. clevelandii, some of these scales are reduced to hairlike structures, which help distinguish it from the closely related M. covillei. It is usually found growing on exposed rock, particularly igneous rock.
Myriopteris gracilis, formerly known as Cheilanthes feei, is a species of lip fern known by the common name slender lip fern or Fee's lip fern.
Myriopteris cooperae, formerly Cheilanthes cooperae, is a species of lip fern known by the common name Mrs. Cooper's lip fern, or simply Cooper's lip fern.
Myriopteris gracillima, formerly known as Cheilanthes gracillima, is a species of lip fern known by the common name lace lip fern. It is native to western North America, where it grows in rocky habitat from British Columbia to California to Montana.
Myriopteris intertexta, formerly Cheilanthes intertexta, is a species of lip fern known by the common name coastal lip fern. It is native to montane California and western Nevada, Oregon east of the Cascades, and with a disjunct population in central Utah. It grows in dry rocky habitats in sun, typically in rock cracks with little or no soil.
Myriopteris tomentosa, formerly known as Cheilanthes tomentosa, is a perennial fern known as woolly lipfern. Woolly lipfern is native to the southern United States, from Pennsylvania to Arizona and Georgia, and Mexico.
Myriopteris, commonly known as the lip ferns, is a genus of cheilanthoid ferns. Like other cheilanthoids, they are ferns of dry habitats, reproducing both sexually and apogamously. Many species have leaves divided into a large number of small, bead-like segments, the probable inspiration for the generic name. Hairs and/or scales are often present on both the upper and lower surfaces of the leaf, and their presence and appearance are useful in distinguishing between species. The genus is most diverse in Mexico, but species are found from southwestern Canada south to southern Chile, and one species is endemic to southern Africa.
Myriopteris alabamensis, the Alabama lip fern, is a moderately-sized fern of the United States and Mexico, a member of the family Pteridaceae. Unlike many members of its genus, its leaves have a few hairs on upper and lower surfaces, or lack them entirely. One of the cheilanthoid ferns, it was usually classified in the genus Cheilanthes as Cheilanthes alabamensis until 2013, when the genus Myriopteris was again recognized as separate from Cheilanthes. It typically grows in shade on limestone outcrops.
Myriopteris aemula, the Texas lip fern or rival lip fern, is a moderately-sized fern of Texas and Mexico, a member of the family Pteridaceae. Unlike many members of its genus, its leaves have a few hairs on upper and lower surfaces, or lack them entirely. One of the cheilanthoid ferns, it was usually classified in the genus Cheilanthes as Cheilanthes aemula until 2013, when the genus Myriopteris was again recognized as separate from Cheilanthes. It typically grows on limestone rock.
Myriopteris yatskievychiana is a small fern native to Mexico, a member of the family Pteridaceae. It is known only from a single collection in the Mexican state of Sonora. While superficially similar to golden lip fern, a widespread species in Mexico, differences in its coating of hairs and its small size make it distinctive. One of the cheilanthoid ferns, it was usually classified in the genus Cheilanthes until 2013, when the genus Myriopteris was again recognized as separate from Cheilanthes.
Myriopteris allosuroides is a moderately-sized fern of Mexico, a member of the family Pteridaceae. Unlike many members of its genus, its rachides are grooved on the upper surface and largely free of hairs or scales. One of the cheilanthoid ferns, it was usually classified in the genera Cheilanthes or Pellaea until 2013, when the genus Myriopteris was again recognized as separate from Cheilanthes. It typically grows on dry, rocky slopes over acidic, particularly basaltic, rock.
Myriopteris aurea, the golden lip fern or Bonaire lip fern, is a moderately-sized fern native to the Americas, a member of the family Pteridaceae. Unlike many members of its genus, its leaf is only modestly dissected into lobed leaflets (pinnae), which are hairy both above and below. One of the cheilanthoid ferns, until 2013 it was classified in the genus Cheilanthes as Cheilanthes bonariensis, when the genus Myriopteris was again recognized as separate from Cheilanthes. It typically grows on dry, rocky slopes, and ranges from Mexico, where it is common and widespread, and the southwestern United States south and east through Central and South America as far as Chile and Argentina.
Myriopteris rawsonii, formerly known as Cheilanthes rawsonii, is a perennial fern native to Namaqualand in Southern Africa. Like many other cheilanthoid ferns, it is adapted to dry conditions, bearing a thick layer of pale hairs on the underside of its pinnate-pinnatifid leaves. It is the only African representative of its clade of cheilanthoids, the otherwise American genus Myriopteris. It spends much of the year in a dried-out, dormant state, rehydrating and putting out new growth during winter rains. Its name honors the botanist and civil servant Sir Rawson W. Rawson.
Myriopteris fendleri, formerly known as Cheilanthes fendleri, is a species of fern in the Pteridaceae family with the common name Fendler's lip fern. It is native to the southwest United States and northern Mexico.
Myriopteris lindheimeri, formerly known as Cheilanthes lindheimeri, is a species of fern in the Pteridaceae family with the common name fairy swords.
Myriopteris yavapensis, formerly known as Cheilanthes yavapensis, is a species of cheilanthoid fern with the common name Yavapai lip fern native to the southwest United States.
Myriopteris wrightii, formerly known as Cheilanthes wrightii, is a species of cheilanthoid fern with the common name Wright's lipfern. It is native to the southwestern United States and northern Mexico.
Myriopteris chipinquensis is a fern endemic to Mexico, a member of the family Pteridaceae. One of the cheilanthoid ferns, it was classified in the genus Cheilanthes until 2013, when the genus Myriopteris was again recognized as separate from Cheilanthes. It grows in oak-pine forests of the Sierra Madre Oriental, often with the very similar and closely related Myriopteris tomentosa.