Vespoidea is a superfamily of wasps in the order Hymenoptera, although older taxonomic schemes may vary in this categorization, particularly in whether to recognize the superfamilies Scolioidea or Formicoidea. Vespoidea includes wasps with a large variety of lifestyles: eusocial, social, and solitary habits, predators, scavengers, parasitoids, and some herbivores.
Myrmeciinae is a subfamily of the Formicidae, ants once found worldwide but now restricted to Australia and New Caledonia. This subfamily is one of several ant subfamilies which possess gamergates, female worker ants which are able to mate and reproduce, thus sustaining the colony after the loss of the queen. The Myrmeciinae subfamily was formerly composed of only one genus, Myrmecia, but the subfamily was redescribed by Ward & Brady in 2003 to include two tribes and four genera: An additional three genera, one form genus, and 9 species were described in 2006 from the Early Eocene of Denmark, Canada, and Washington. Subsequently an additional fossil genus was moved from the family Rhopalosomatidae in 2018, and a new genus was described in 2021.
Rhopalosomatidae is a family of Hymenoptera containing about 68 extant species in four genera that are found worldwide. Three fossil genera are known.
Horntail or wood wasp are any of the 150 non-social species of the hymenopteran family Siricidae, a type of wood-eating sawfly. The common name "horntail" derives from the stout, spine-like structure at the end of the adult's abdomen, which is used to pierce the host's bark to allow the eggs to be inserted into the wood. A typical adult horntail is brown, blue, or black with yellow or red parts, and may often reach up to 4 cm (1.6 in) long. The pigeon horntail can grow up to 5 cm (2.0 in) long, among the longest of all Hymenoptera.
Pamphiliidae is a small wasp family within Symphyta, containing some 200 species from the temperate regions of North America and Eurasia. The larvae feed on plants, using silk to build webs or tents, or to roll leaves into tubes in which they feed, thus earning them the common names leaf-rolling sawflies or web-spinning sawflies. Some species are gregarious and the larvae live in large groups. Fossils of Pamphiliidae have been dated to the Jurassic period.
Pelecinidae is a family of parasitic wasps in the Proctotrupoidea. It contains only one living genus, Pelecinus, with three species known from the Americas. The earliest fossil species are known from the Jurassic, and the group was highly diverse during the Cretaceous. Members of Pelecinus are parasitic on larval beetles, flies, green lacewings, and sawflies.
The Stephanidae, sometimes called crown wasps, are a family of parasitoid wasps. They are the only living members of the superfamily Stephanoidea. Stephanidae has at least 345 living species in 11 genera. The family is considered cosmopolitan in distribution, with the highest species concentrations in subtropical and moderate climate zones. Stephanidae also contain four extinct genera described from both compression fossils and inclusions in amber.
Ulteramus is an extinct genus of parasitic wasp in the sawfly family Pamphiliidae. The genus is solely known from an Eocene fossil found in North America. At the time of its description the new genus was composed of a single species, Ulteramus republicensis.
The Klondike Mountain Formation is an Early Eocene (Ypresian) geological formation located in the northeast central area of Washington state. The formation, named for the type location designated in 1962, Klondike Mountain north of Republic, Washington, is composed of volcanic rocks in the upper unit and volcanics plus lacustrine (lakebed) sedimentation in which a lagerstätte with exceptionally well-preserved plant and insect fossils has been found, along with fossil epithermal hot springs.
Ypresiomyrma is an extinct genus of ants in the subfamily Myrmeciinae that was described in 2006. There are four species described; one species is from the Isle of Fur in Denmark, two are from the McAbee Fossil Beds in British Columbia, Canada, and the fourth from the Bol’shaya Svetlovodnaya fossil site in Russia. The queens of this genus are large, the mandibles are elongated and the eyes are well developed; a stinger is also present. The behaviour of these ants would have been similar to that of extant Myrmeciinae ants, such as solitary foraging for arthropod prey and never leaving pheromone trails. The alates were poor flyers due to their size, and birds and animals most likely preyed on these ants. Ypresiomyrma is not assigned to any tribe, and is instead generally regarded as incertae sedis within Myrmeciinae. However, some authors believe Ypresiomyrma should be assigned as incertae sedis within Formicidae.
Macabeemyrma is an extinct genus of bulldog ants in the subfamily Myrmeciinae containing the single species Macabeemyrma ovata, described in 2006 from Ypresian stage deposits of British Columbia, Canada. Only a single specimen is known; a holotype queen found preserved as a compression fossil. The specimen had no wings and small portions of its legs and eyes were faintly preserved. It was a large ant, reaching 25 millimetres (0.98 in) in length. This ants' behaviour would have been similar to that of extant Myrmeciinae ants, such as foraging singly in search for arthropod prey and nesting in soil or in trees. Macabeemyrma shows similarities to extinct ants in the genus Ypresiomyrma, and to the living Nothomyrmecia macrops, but has not been conclusively assigned to any tribe, instead generally regarded as incertae sedis within Myrmeciinae. However, the sole specimen lacks definitive traits, and its classification in Myrmeciinae, and even its identity as an ant, has been challenged.
The McAbee Fossil Beds is a Heritage Site that protects an Eocene Epoch fossil locality east of Cache Creek, British Columbia, Canada, just north of and visible from Provincial Highway 97 / the Trans-Canada Highway. The McAbee Fossil Beds, comprising 548.23 hectares, were officially designated a Provincial Heritage Site under British Columbia's Heritage Conservation Act on July 19, 2012. The site is part of an old lake bed which was deposited about 52 million years ago and is internationally recognised for the diversity of plant, insect, and fish fossils found there. Similar fossil beds in Eocene lake sediments, also known for their well preserved plant, insect and fish fossils, are found at Driftwood Canyon Provincial Park near Smithers in northern British Columbia, on the Horsefly River near Quesnel in central British Columbia, and at Republic in Washington, United States. The Princeton Chert fossil beds in southern British Columbia are also Eocene, but primarily preserve an aquatic plant community. A 2016 review of the early Eocene fossil sites from the interior of British Columbia discusses the history of paleobotanical research at McAbee, the Princeton Chert, Driftwood Canyon, and related Eocene fossil sites such as at Republic.
The Coldwater Beds are a geologic formation of the Okanagan Highlands in British Columbia, Canada. They preserve fossils dating back to the Ypresian stage of the Eocene period, or Wasatchian in the NALMA classification.
Aphaenogaster longaeva is an extinct species of ant in formicid subfamily Myrmicinae known from a solitary Eocene or Oligocene fossil found in North America. A. longaeva was one of five insect species described by the paleoentomologist Samuel Hubbard Scudder in an 1877 paper.
This list of fossil arthropods described in 2009 is a list of new taxa of trilobites, fossil insects, crustaceans, arachnids and other fossil arthropods that have been described during the year 2009, as well as other significant discoveries and events related to arthropod paleontology that occurred.
Cuspilongus is an extinct genus of parasitic wasp in the sawfly family Cephidae. At the time of its description the new genus comprised a single species named Cuspilongus cachecreekensis, known from an Eocene fossil found in North America. Subsequently, Kopylov & Rasnitsyn (2016) transferred the Early Cretaceous (Aptian) species Mesocephus ghilarovi Rasnitsyn (1988), known from the Bon-Tsagan locality in Mongolia, to the genus Cuspilongus as well.
Aptenoperissus is a genus of extinct wasp with eight described species, placed into the monotypic family Aptenoperissidae. The type species Aptenoperissus burmanicus resembles a mix between a grasshopper, an ant, and a wasp. It was described by a group of researchers from Oregon State University in a paper released online in October 2016. The piece of 100 million year old Burmese amber that it was preserved in was found in the Hukawng Valley of Myanmar in Southern Asia. A new family, Aptenoperissidae, was described to accommodate this insect. Species of Aptenoperissus were wingless, with a strong stinger. The creature had long legs making it perfect for jumping higher than most insects. Subsequently additional species were described from the Myanmar amber: A. amabilis, A. delicatus, A. formosus, A. etius, A. magnifemoris, A. pusillus and A. zonalis. Initially placed in Ceraphronoidea, later studies placed it in Stephanoidea.
Embolemidae is a family of small solitary parasitoid wasps with around 70 species in 2 genera distributed around the world. The few species whose biology is known are parasites on planthopper nymphs of the families Achilidae and Cixiidae. There is debate regarding the status of the genus named Ampulicomorpha by Ashmead in 1893, generally considered now to be a junior synonym of Embolemus (e.g.,), though some authorities dispute this (e.g.,)
Burmese amber is fossil resin dating to the early Late Cretaceous Cenomanian age recovered from deposits in the Hukawng Valley of northern Myanmar. It is known for being one of the most diverse Cretaceous age amber paleobiotas, containing rich arthropod fossils, along with uncommon vertebrate fossils and even rare marine inclusions. A mostly complete list of all taxa described up until 2018 can be found in Ross 2018; its supplement Ross 2019b covers most of 2019.
Klondikia is an extinct hymenopteran genus in the ant family Formicidae with a single described species Klondikia whiteae. The species is solely known from the Early Eocene sediments exposed in northeast Washington state, United States. The genus is currently not placed into any ant subfamily, being treated as incertae sedis.
