Ptolemaiida | |
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Lower jaw of Ptolemaia lyonsi | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Superorder: | Afrotheria |
Grandorder: | Afroinsectiphilia |
Order: | † Ptolemaiida Simons & Bown, 1995 |
Families and genera | |
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Ptolemaiida is a taxon of wolf-sized afrothere mammals that lived in northern and eastern Africa during the Paleogene. The oldest fossils are from the latest Eocene strata of the Jebel Qatrani Formation, near the Fayum oasis in Egypt. [1] A tooth is known from an Oligocene-aged stratum in Angola, [2] and Miocene specimens (of Kelba ) are known from Kenya and Uganda. [3]
The origin of the Ptolemaiida is obscure, and debated. The type species was originally thought to be a primate, but, later, when elongated skulls with long canines of Ptolemaia and Qarunavus were found, they were then thought to be hyaenodontids, or giant, carnivorous relatives of the pantolestids Palaeosinopa , and of modern shrews [4] The family Ptolemaiidae was elevated to order level in 1995, [1] although some experts later placed the Ptolemaiidae within the pantolestids. [5]
Recently, Ptolemaiida has been placed within Afrotheria [6] on the basis of paleobiology, as the taxon was endemic to Africa, and because of some similarities in the anatomical features of the skull in common with aardvarks. [7] [8] It is currently unclear if they form a sister taxon to Tubulidentata or are a paraphyletic sequence leading to them. Regardless, their close relation may offer the possibility for true dental synapomorphies in Afroinsectiphilia. [7] [8] A posterior study on ptolemaiidan dentitions further reinforces these results. [9]
Recently, there has been a reconsideration of the ptolemaiidan diet, and possible behavior, as wear on the teeth suggest that it crushed hard or abrasive food, and that the teeth had little or no shearing ability. Even so, some sources still refer to them as being gigantic, carnivorous shrews. [10]
A tenrec is a mammal belonging to any species within the afrotherian family Tenrecidae, which is endemic to Madagascar. Tenrecs are a very diverse group; as a result of convergent evolution, some resemble hedgehogs, shrews, opossums, rats, and mice. They occupy aquatic, arboreal, terrestrial, and fossorial environments. Some of these species, including the greater hedgehog tenrec, can be found in the Madagascar dry deciduous forests. However, the speciation rate in this group has been higher in humid forests.
Afrotheria is a superorder of placental mammals, the living members of which belong to groups that are either currently living in Africa or of African origin: golden moles, elephant shrews, otter shrews, tenrecs, aardvarks, hyraxes, elephants, sea cows, and several extinct clades. Most groups of afrotheres share little or no superficial resemblance, and their similarities have only become known in recent times because of genetics and molecular studies. Many afrothere groups are found mostly or exclusively in Africa, reflecting the fact that Africa was an island continent from the Cretaceous until the early Miocene around 20 million years ago, when Afro-Arabia collided with Eurasia.
Paenungulata is a clade of "sub-ungulates", which groups three extant mammal orders: Proboscidea, Sirenia, and Hyracoidea (hyraxes). At least two more possible orders are known only as fossils, namely Embrithopoda and Desmostylia.
Orycteropodidae is a family of afrotherian mammals. Although there are many fossil species, the only species surviving today is the aardvark, Orycteropus afer. Orycteropodidae is recognized as the only family within the order Tubulidentata, so the two are effectively synonyms.
Aegyptopithecus is an early fossil catarrhine that predates the divergence between hominoids (apes) and cercopithecids. It is known from a single species, Aegyptopithecus zeuxis, which lived around 38-29.5 million years ago in the early part of the Oligocene epoch. It likely resembled modern-day New World monkeys, and was about the same size as a modern howler monkey, which is about 56 to 92 cm long. Aegyptopithecus fossils have been found in the Jebel Qatrani Formation of modern-day Egypt. Aegyptopithecus is believed to be a stem-catarrhine, a crucial link between Eocene and Miocene fossils.
Cimolesta is an extinct order of non-placental eutherian mammals. Cimolestans had a wide variety of body shapes, dentition and lifestyles, though the majority of them were small to medium-sized general mammals that bore superficial resemblances to rodents, lagomorphs, mustelids, and marsupials.
The Afroinsectiphilia is a clade that has been proposed based on the results of recent molecular phylogenetic studies. Many of the taxa within it were once regarded as part of the order Insectivora, but Insectivora is now considered to be polyphyletic and obsolete. This proposed classification is based on molecular studies only, and there is no morphological evidence for it.
Eogavialis is an extinct genus of eusuchian crocodylomorph, usually regarded as a gavialoid crocodylian. It superficially resembles Tomistoma schlegelii, the extant false gharial, and consequently material from the genus was originally referred to Tomistoma. Indeed, it was not until 1982 that the name Eogavialis was constructed after it was realised that the specimens were from a more basal form.
The Jebel Qatrani Formation is a geologic formation located in the Faiyum Governorate of central Egypt. It is exposed between the Jebel Qatrani escarpment and the Qasr el Sagha escarpment, north of Birket Qarun lake near Faiyum. The formation conformably overlies the Qasr el Sagha Formation and is topped by the Widan el Faras Basalt. The age of the formation has been subject to debate, but the most recent research indicates that it covers both the latest parts of the Eocene and the Early Oligocene, spanning over the boundary between these two time periods.
Parapithecidae is an now extinct family of primates which lived in the Eocene and Oligocene periods in Egypt. Eocene fossils from Myanmar are sometimes included in the family in addition. They showed certain similarities in dentition to Condylarthra, but had short faces and jaws shaped like those of tarsiers. They are part of the superfamily Parapithecoidea, perhaps equally related to Ceboidea and Cercopithecoidea plus Hominoidea - but the placement of Parapithecoidea is substantially uncertain.
Plesiopithecus is an extinct genus of early strepsirrhine primate from the late Eocene.
Kelba quadeemae is an extinct species of ptolemaiidan mammal, the sole species of the family Kelbidae, known from the Lower Miocene of East Africa. The genus name Kelba derives from the Arabic الكلب meaning "dog", and the specific name quadeemae from the Arabic quadeem, meaning "ancient". Kelba is only known from a partial skull and teeth, but estimated to have been, similar in size to a coyote but more heavily built. The teeth are rather unspecialized, suggesting a wide and varied diet, and show wear suggesting its diet included abrasive material.
Ptolemaia is a genus of extinct Afrotherian mammals from the Oligocene of East Africa. The genus and type species, P. lyonsi, was described by Henry Fairfield Osborn in 1908 from the Jebel Qatrani Formation of Egypts' Fayum Depression. The genus name alludes to the Ptolemaic dynasty of Ancient Greece which ruled over the Egyptian region where Ptolemaia was discovered, while the specific epithet lyonsi honors H. G. Lyons, then director of the Egyptian Geological Survey. A second species, P. grangeri, was described in 1987, and named after the early 20th century paleontologist Walter W. Granger. Fossils of P. grangeri are also known from Kenya.
Widanelfarasia is an extinct genus of placental mammals known from the Late Eocene Jebel Qatrani Formation of Egypt. Two species are known: W. bowni and the smaller W. rasmusseni. Described in 2000 by E. R. Seiffert and Elwyn L. Simons, Widanelfarasia was initially classified as uncertain position within placentals, but was later placed within the afrosoricidan suborder Tenrecomorpha. The genus name derives from Widan el-Faras, two prominent hills in the area where the fossils were recovered.
Hyaenodonta is an extinct order of hypercarnivorous placental mammals of clade Pan-Carnivora from mirorder Ferae. Hyaenodonts were important mammalian predators that arose during the early Paleocene in Europe and persisted well into the late Miocene.
Propliopithecoidea is a superfamily of catarrhine primates that inhabited Africa and the Arabian Peninsula during the Early Oligocene about 32 to 29 million years ago. Fossils have been found in Egypt, Oman and Angola. They are one of the earliest known families of catarrhines. They have a number of features in common with extant catarrhines, but also features that are primitive and not found in later catarrhine families.
Parapithecoidea is an extinct superfamily of primates which lived in the Eocene and Oligocene periods in Egypt. In some classifications all Parapithecoidea are placed within the family Parapithecidae. Seiffert et al. (2010) propose that Parapithecoidea arose during the Bartonian, with a split between Biretia and the Parapithecidae occurring early in the Priabonian.
Metapterodon is an extinct genus of hyainailourid hyaenodonts of the subfamily Hyainailourinae, that lived in Africa during the early Oligocene to early Miocene. Fossils of Metapterodon were recovered from the Egypt, Uganda, Elisabeth Bay Formation in Namibia, and Rusinga Island and Karungu in Kenya.
Hesham Sallam is an Egyptian paleontologist and the founder of the Mansoura University Vertebrate Paleontology Center (MUVP-C), the first vertebrate paleontology program in the Middle East. He works as an associate professor at the American University in Cairo and Mansoura University. Sallam led the discovery and description of Mansourasaurus shahinae, a species of sauropod dinosaur from Egypt, which has improved understanding of the prehistory of Africa during the latest Cretaceous period. His work has helped popularize paleontology in Egypt.
Altacreodus is an extinct genus of eutherian mammals. Fossils have been found in North America where they first appeared during the Late Cretaceous, and they died out prior to the start of the Paleocene. It is possibly one of the earliest known placental mammals in the fossil record.
In addition to these large mammals, the small mammal fauna of the Fayûm beds of Africa include many different insectivores (including one, Ptolemaia, that became a wolf-sized predator) ....