Rhombocorniculum

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Rhombocorniculum
Temporal range: Cambrian Stage 2–Cambrian Stage 3 [1]
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Scientific classification Red Pencil Icon.png
Kingdom: Animalia
Family: Hallucigeniidae
Genus:Rhombocorniculum
Walliser, 1958 [2]

Rhombocorniculum is a species of small shelly fossil comprising twisted ornamented cones. It has been described from the Comely limestone and elsewhere. R. cancellatum straddles the Atdabanian/Botomian boundary. [1] The structure of its inner layer suggests that its phosphatic fibres formed within a flexible organic matrix. [3]

Contents

Taxonomy

Three species are recognized — in stratigraphic succession: R. insolutum, R. cancellatum (=R. walliseri), and R. spinosus (=Rushtonites spinosus). [4] Landing (1995) refers R. insolutum to the strictocorniculids, along with Rushtonites . [3] Hinz (1987) considers insolutum to fall within the variability seen in cancellatum.

Affinity

Based on details of the ornament and construction, Rhombocorniculum is interpreted as the spines of a Hallucigenia -like lobopodian worm. [5]

<i>Hallucigenia</i> genus of Cambrian animals

Hallucigenia is a genus of Cambrian xenusiids known from articulated fossils in Burgess Shale-type deposits in Canada and China, and from isolated spines around the world. The generic name reflects the type species' unusual appearance and eccentric history of study; when it was erected as a genus, H. sparsa was reconstructed upside down and back to front. Hallucigenia is now recognized as a "lobopodian worm". It is considered by some to represent an early ancestor of the living velvet worms, although other researchers favour a relationship closer to arthropods.

See also

Related Research Articles

The Cambrian Period was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 55.6 million years from the end of the preceding Ediacaran Period 541 million years ago (mya) to the beginning of the Ordovician Period 485.4 mya. Its subdivisions, and its base, are somewhat in flux. The period was established by Adam Sedgwick, who named it after Cambria, the Latin name of Wales, where Britain's Cambrian rocks are best exposed. The Cambrian is unique in its unusually high proportion of lagerstätte sedimentary deposits, sites of exceptional preservation where "soft" parts of organisms are preserved as well as their more resistant shells. As a result, our understanding of the Cambrian biology surpasses that of some later periods.

The Ediacaran Period, spans 94 million years from the end of the Cryogenian Period 635 million years ago (Mya), to the beginning of the Cambrian Period 541 Mya. It marks the end of the Proterozoic Eon, and the beginning of the Phanerozoic Eon. It is named after the Ediacara Hills of South Australia.

Global Boundary Stratotype Section and Point

A Global Boundary Stratotype Section and Point, abbreviated GSSP, is an internationally agreed upon reference point on a stratigraphic section which defines the lower boundary of a stage on the geologic time scale. The effort to define GSSPs is conducted by the International Commission on Stratigraphy, a part of the International Union of Geological Sciences. Most, but not all, GSSPs are based on paleontological changes. Hence GSSPs are usually described in terms of transitions between different faunal stages, though far more faunal stages have been described than GSSPs. The GSSP definition effort commenced in 1977. As of 2012, 64 of the 101 stages that need a GSSP have been formally defined.

Lobopodia group of animals

The lobopodians, members of the informal group Lobopodia are worm-like taxa with stubby legs.

<i>Microdictyon</i> genus of primitive animal (fossil)

Microdictyon is an extinct "armored worm" coated with net-like scleritic scales, known from the Early Cambrian Maotianshan shale of Yunnan China. Microdictyon is sometimes included in a somewhat ill-defined Phylum – Lobopodia – that includes several other odd worm-like and segmented free-swimming animals that do not appear to be arthropods or worms. The phylum includes Microdictyon, Onychodictyon, Cardiodictyon, Luolishania, Paucipodia, as well as the Anomalocaridids. The isolated sclerites of Microdictyon are known from other Lower Cambrian deposits. Microdictyon sclerites appear to have moulted; one sclerite seems to have been preserved during ecdysis.

Halkieriid Family of molluscs

The halkieriids are a group of fossil organisms from the Lower to Middle Cambrian. Their eponymous genus is Halkieria, which has been found on almost every continent in Lower to Mid Cambrian deposits, forming a large component of the small shelly fossil assemblages. The best known species is Halkieria evangelista, from the North Greenland Sirius Passet Lagerstätte, in which complete specimens were collected on an expedition in 1989. The fossils were described by Simon Conway Morris and John Peel in a short paper in 1990 in the journal Nature. Later a more thorough description was undertaken in 1995 in the journal Philosophical Transactions of the Royal Society of London and wider evolutionary implications were posed.

The small shelly fauna, small shelly fossils (SSF), or early skeletal fossils (ESF) are mineralized fossils, many only a few millimetres long, with a nearly continuous record from the latest stages of the Ediacaran to the end of the Early Cambrian Period. They are very diverse, and there is no formal definition of "small shelly fauna" or "small shelly fossils". Almost all are from earlier rocks than more familiar fossils such as trilobites. Since most SSFs were preserved by being covered quickly with phosphate and this method of preservation is mainly limited to the Late Ediacaran and Early Cambrian periods, the animals that made them may actually have arisen earlier and persisted after this time span.

Stratigraphy of the Cambrian

The Stratigraphy of the Cambrian period currently has several schemes used for ordering geologic formations from the period. The International Commission on Stratigraphy−ICS scheme has set a stratotype section for the base of the Cambrian, dated quite accurately to 541 ± 1.0 million years ago. Russian and Chinese scientists have developed a different scheme.

Mount Cap formation

The Mount Cap Formation is a geologic formation exposed in the Mackenzie Mountains, northern Canada. It was deposited in a shallow shelf setting in the late Early Cambrian, and contains an array of Burgess Shale-type microfossils that have been recovered by acid maceration.

Turcutheca is a Tommotian genus of shelly fossil whose affinities are uncertain, generally considered as an orthothecid hyolith (which would make it a brachiopod, but also resembling the ellesmeroceratids.

Mongolitubulus is a form genus encapsulating a range of ornamented conical small shelly fossils of the Cambrian period. It is potentially synonymous with Rushtonites, Tubuterium and certain species of Rhombocorniculum, and owing to the similarity of the genera, they are all dealt with herein. Organisms that bore Mongolitubulus-like projections include trilobites, bradoriid arthropods and hallucigeniid lobopodians.

Martin David Brasier FGS, FLS was an English palaeobiologist and astrobiologist known for his conceptual analysis of microfossils and evolution in the Precambrian and Cambrian.

Terreneuvian first epoch of the Cambrian Period

The Terreneuvian is the lowermost and oldest series of the Cambrian geological system. Its base is defined by the first appearance datum of the trace fossil Treptichnus pedum around 541 million years ago. Its top is defined as the first appearance of trilobites in the stratigraphic record around 521 million years ago. This series was formally ratified by the International Commission on Stratigraphy in 2012.

Fortunian first age of the Terreneuvian Epoch

The Fortunian stage marks the beginning of the Phanerozoic eon, the Paleozoic era, and the Cambrian period. It is the first of the two stages of the Terreneuvian series. Its base is defined as the first appearance of the trace fossil Treptichnus pedum 541 million years ago. The top of the Fortunian which is the base of the Stage 2 of the Cambrian has not been formally defined yet, but will correspond to the appearance of an Archeocyatha species or "Small shelly fossils" approximately 529 million years ago.

Chapel Island Formation

The Chapel Island Formation is a sedimentary formation from the Burin Peninsula, Newfoundland, Canada. It is a succession of siliciclastic deposits, over 1,000 metres (3,300 ft) thick, that were deposited during the latest Ediacaran and earliest Cambrian.

Quadratapora is a genus of lobopodian known only from its biomineralized dorsal plates, which somewhat resemble those of Microdictyon. Its fossils date to the Tommotian, representing the earliest record of lobopodians.

Random Formation

The Random Formation is a rock unit in Newfoundland dating to the early Cambrian period, dominated by tidal quartz arenites deposited in a near-shore environment, but also incorporating intertidal and open-shelf deposits, including glauconitic and mud-cracked mudstones, and red channel sandstones. It was deposited quickly and is approximately 175 m thick. The Blue Pinion Formation was originally recognized as a separate formation, but is now interpreted as an expression of the Random Formation.

Circothecidae are a family of Cambrian problematica, sometimes attributed to the Hyolitha, though some authors suggest that they're definitely not.

References

  1. 1 2 Brasier, M. D. (1986). "The succession of small shelly fossils (especially conoidal microfossils) from English Precambrian–Cambrian boundary beds". Geological Magazine. 123 (3): 237. doi:10.1017/S0016756800034737.
  2. Otto H. Walliser. "Rhombocorniculum comleyense n. gen., n. sp". Paläontologische Zeitschrift. 32 (3–4): 176–180. doi:10.1007/BF02989029.
  3. 1 2 Landing, E. (May 1995). "Upper Placentian-Branchian Series of Mainland Nova Scotia (Middle-Upper Lower Cambrian): Faunas, Paleoenvironments, and Stratigraphic Revision". Journal of Paleontology. Paleontological Society. 69 (3): 475–495. JSTOR   1306322.
  4. Brasier, M. D. (1989). Towards a biostratigraphy of the earliest skeletal biotas. In J. W. Cowie & M. D. Brasier (Eds.), The Precambrian-Cambrian boundary (pp. 117–165). Oxford: Clarendon Press.
  5. Caron, J. -B.; Smith, M. R.; Harvey, T. H. P. (2013). "Beyond the Burgess Shale: Cambrian microfossils track the rise and fall of hallucigeniid lobopodians". Proceedings of the Royal Society B: Biological Sciences. 280 (1767): 20131613. doi:10.1098/rspb.2013.1613. PMC   3735267 Lock-green.svg. PMID   23902914.