Salsoloideae | |
---|---|
Salsola oppositifolia , in fruit | |
Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Angiosperms |
Clade: | Eudicots |
Order: | Caryophyllales |
Family: | Amaranthaceae |
Subfamily: | Salsoloideae Raf. |
Genera | |
About 35 genera, see text |
The Salsoloideae are a subfamily of the Amaranthaceae, formerly in family Chenopodiaceae.
These are herbs, subshrubs, shrubs and some trees. Stems and leaves are often succulent. The ovary contains a spiral embryo. In most genera, scarious wings develop at the outside of the fruiting perianth, allowing for dispersal by the wind (anemochory). In tribe Caroxyleae, the stamens have vesiculose anther appendages, discolor with anthers, that probably play a role for insect pollination. In tribe Salsoleae the anther appendages are absent or small and inconspicuous.
The area with most species (center of diversity) are the deserts and semideserts of Central-Asia and the Middle East. Distribution of the subfamily extends to the Mediterranean, to Middle-Europe, north and south Africa, and Australia, some species have also been introduced to America. Many species grow in dry habitats (xerophytes) or tolerate salty soils (halophytes), some are ruderals.
Salsoloideae are C4 plants (with a few exceptions in tribe Salsoleae). Tribe Caroxyleae is exclusively of the NAD-malic enzyme C4 subtype. Most Salsoleae also use the NADP-malic enzyme. [1]
The type genus of subfamily Salsoloideae is Salsola L. According to recent research, the former classification did not reflect the phylogenetic relationship, so that the subfamily had to be reclassified; it is now split in two tribes, Caroxyleae and Salsoleae (in the strict sense). [2] [3] The tribe Camphorosmeae is now treated in its own subfamily, Camphorosmoideae. [4]
Caroxyleae Akhani & E. H. Roalson (as "Caroxyloneae") [2]
Amaranthaceae is a family of flowering plants commonly known as the amaranth family, in reference to its type genus Amaranthus. It includes the former goosefoot family Chenopodiaceae and contains about 165 genera and 2,040 species, making it the most species-rich lineage within its parent order, Caryophyllales.
Halogeton is a plant genus of the family Amaranthaceae. The genus name, Halogeton, derives from the Greek words for "salt" and for "neighbor."
Suaeda is a genus of plants also known as seepweeds and sea-blites. Most species are confined to saline or alkaline soil habitats, such as coastal salt-flats and tidal wetlands. Many species have thick, succulent leaves, a characteristic seen in various plant genera that thrive in salty habitats.
Salsola is a genus of the subfamily Salsoloideae in the family Amaranthaceae. The genus sensu stricto is distributed in central and southwestern Asia, North Africa, and the Mediterranean. A common name of various members of this genus and related genera is saltwort, for their salt tolerance. The genus name Salsola is from the Latin salsus, meaning "salty".
Halothamnus is a genus in the subfamily Salsoloideae of the family Amaranthaceae. The scientific name means saltbush, from the Greek ἅλς (hals) "salt" and θαμνος (thamnos) "bush". This refers either to salty habitats or to the accumulation of salt in the plants. The genus is distributed from Southwest and Central Asia to the Arabian peninsula and East Africa.
Halothamnus glaucus is a species of the plant genus Halothamnus, that belongs to the subfamily Salsoloideae of the family Amaranthaceae,. It occurs in Western and Central Asia.
Halothamnus auriculus is a species of the plant genus Halothamnus, that belongs to the subfamily Salsoloideae of the family Amaranthaceae,. It occurs in Western and Middle Asia.
Bassia is a genus of flowering plants in the family Amaranthaceae. They are distributed in the western Mediterranean to eastern Asia. Some occur outside their native ranges as introduced species.
Haloxylon is a genus of shrubs or small trees, belonging to the plant family Amaranthaceae. Haloxylon and its species are known by the common name saxaul. "Saksaul" is a common Turkic word that entered Russian through Kazakh.
Camphorosmeae is a species-rich tribe of the Amaranthaceae, formerly Chenopodiaceae, with 20 genera and about 179 species. It is classified as a single tribe of subfamily Camphorosmoideae.
The Polycnemoideae are a small subfamily of plants in the family Amaranthaceae, representing a basal evolutionary lineage. The few relictual species are distributed in Eurasia and North Africa, North America, and Australia.
The Suaedoideae are a subfamily of plants in the family Amaranthaceae.
Kali was a genus of plants in the subfamily Salsoloideae in the family Amaranthaceae, that has now been subsumed into the genus Salsola.
Anabasis is a genus of plants in the subfamily Salsoloideae of the family Amaranthaceae. It is distributed in southern Europe, North Africa, and Asia.
Caroxylon imbricatum, synonym Salsola imbricata, is a small species of shrub in the family Amaranthaceae. It grows in deserts and arid regions of north Africa, the Arabian Peninsula and southwestern Asia.
Caroxylon is a genus of shrubby flowering plants in the family Amaranthaceae, found in drier areas of the Old World, including southern Africa, Madagascar, northern Africa, Mediterranean islands of Europe, the Canary Islands, Socotra, Ukraine, Russia, western Asia, Central Asia, India, western and northern China, and Mongolia.
Horaninovia is a genus of flowering plants belonging to the family Amaranthaceae. It also belongs to the tribe Salsoleae as well as in the subfamily Salsoloideae.
Hammada is a genus of flowering plants belonging to the family Amaranthaceae. It is also in the Salsoloideae subfamily. Although it is a very unclear and unsorted genus with many species that have later been classed as synonyms.
Akhania is a genus of flowering plants in the amaranth family, Amaranthaceae. It includes three species native to the eastern Mediterranean and western and central Asia, ranging from Greece through western Asia to the Caucasus, Kazakhstan, and Afghanistan.