Tarachoptera

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Tarachoptera
Temporal range: Cenomanian
Tarachocelis microlepidopterella holotype.jpg
Tarachocelis microlepidopterella
Kinitocelis divisinotata holotype Fig29.jpg
Kinitocelis divisinotata
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
(unranked): Amphiesmenoptera
Order: Tarachoptera
Mey, Wichard, Müller and Wang, 2017
Families

Tarachoptera is an extinct order of insects, currently solely known from the mid Cretaceous aged Burmese amber. It belongs to Amphiesmenoptera alongside living Lepidopterans (butterflies and moths) and Trichoptera (caddiesflies), but is outside the clade containing the two groups, with an estimated divergence during the Upper Triassic, and therefore over a 100 million year ghost lineage. It currently contains only one family, the Tarachocelidae, which was named in a publication in 2017 a few months before the order itself was published. [1] As the manuscript was in submission, an additional specimen belonging to a new genus justified creating a separate order. [2] Additional species were described in 2018 [3] and 2020. [4] A notable character is the possession of wing scales like members of Lepidoptera, these were initially suggested to have been evolved in parallel evolution from hairs or setae, [2] but a later study suggested that the scales of all amphiesmenopterans are homologous, sharing a common origin. [5] Their flattened morphology and small size suggest they were adapted to living in small crevices; the morphology of the mouth suggests that they were phytophagous, and ingested small particles, perhaps including pollen grains. [2]

Taxonomy

Related Research Articles

<span class="mw-page-title-main">Amphiesmenoptera</span> Superorder of insects

Amphiesmenoptera is an insect superorder, established by S. G. Kiriakoff, but often credited to Willi Hennig in his revision of insect taxonomy for two sister orders: Lepidoptera and Trichoptera (caddisflies). In 2017, a third fossil order was added to the group, the Tarachoptera.

<span class="mw-page-title-main">Compsocidae</span> Family of booklice

Compsocidae is a family of Psocodea belonging to the suborder Troctomorpha. The family comprises two extant species in two genera, both found in Mesoamerica. Compsocus elegans is found in Mexico and Central America, while Electrentomopsis variegata is found in Mexico. The antennae of each species have 13 or 14 segments. Two extinct genera, Burmacompsocus and Paraelectrentomopsis are known from the Cenomanian aged Burmese amber of Myanmar and Albian aged Spanish amber.

<span class="mw-page-title-main">Ommatidae</span> Family of beetles

The Ommatidae are a family of beetles in the suborder Archostemata. The Ommatidae are considered the extant beetle family that has most ancestral characteristics. There are only seven extant species, confined to Australia and South America. However, the geographical distribution was much wider during the Mesozoic spanning across Eurasia and Australia, suggesting that they were widespread on Pangea. So far, over 26 extinct genera containing over 170 species of these beetles have been described. Three extant genera have been assigned to this family: Omma,Tetraphalerus and Beutelius. The family is considered to be a subfamily of Cupedidae by some authors, but have been found to be more closely related to Micromalthidae in molecular phylogenies. A close relationship with Micromalthidae is supported by several morphological characters, including those of the mandibles and male genitalia. Due to their rarity, their ecology is obscure, it is likely that their larvae feed on deadwood.

<span class="mw-page-title-main">Lepidoptera fossil record</span>

The Lepidoptera fossil record encompasses all butterflies and moths that lived before recorded history. The fossil record for Lepidoptera is lacking in comparison to other winged species, and tending not to be as common as some other insects in the habitats that are most conducive to fossilization, such as lakes and ponds, and their juvenile stage has only the head capsule as a hard part that might be preserved. Yet there are fossils, some preserved in amber and some in very fine sediments. Leaf mines are also seen in fossil leaves, although the interpretation of them is tricky. Putative fossil stem group representatives of Amphiesmenoptera are known from the Triassic.

<span class="mw-page-title-main">Dysoneuridae</span> Extinct family of caddisflies

Dysoneuridae is an extinct family of insect in the order Trichoptera, the caddisflies. The family was first described by I.D. Sukacheva in 1968, and lived from the Middle Jurassic to mid-Cretaceous.

<span class="mw-page-title-main">Alienopteridae</span> Extinct family of cockroaches

Alienopteridae is an extinct family of dictyopterans, known from the Mid-Cretaceous to Eocene. They are noted for their unusual combination of features not found in other dictyopterans.

2018 in paleoentomology is a list of new fossil insect taxa that were described during the year 2018, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.

2019 in paleoentomology is a list of new fossil insect taxa that were described during the year 2019, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.

2017 in paleoentomology is a list of new fossil insect taxa that were described during the year 2017, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.

2020 in paleoentomology is a list of new fossil insect taxa that were described during the year 2020, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.

<span class="mw-page-title-main">Permopsocida</span> Extinct genus of insects

Permopsocida is an extinct order of insects known from the Carboniferous (Moscovian) to the Mid-Cretaceous. It is part of Paraneoptera, alongside bark lice, bugs and thrips. Within Paraneoptera it is considered to be closer to the clade containing bugs and thrips rather than bark lice, with an estimated divergence during the Late Carboniferous. The group was first named as a suborder by Robert John Tillyard in 1926, and was raised to a full order by Huang et al. in 2016. It is currently divided up into three families, Psocidiidae which is known from the Permian to Liassic. Permopsocidae which is only known from the Permian, and Archipsyllidae, which is known from the Late Triassic to mid-Cretaceous (Cenomanian). While most members of the group are known from compression fossils, several members of Archipsyllidae are 3 dimensionally preserved in Burmese amber, which has helped clarify the morphology and phylogenetic position of the group. The morphology of the mouthparts suggests that they were capable of suction feeding and chewing, with preserved angiosperm pollen grains in the gut of Psocorrhyncha suggesting that at least some members of the group were pollenivorous.

2015 in paleoentomology is a list of new fossil insect taxa that were described during the year 2016, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.

Burmese amber is fossil resin dating to the early Late Cretaceous Cenomanian age recovered from deposits in the Hukawng Valley of northern Myanmar. It is known for being one of the most diverse Cretaceous age amber paleobiotas, containing rich arthropod fossils, along with uncommon vertebrate fossils and even rare marine inclusions. A mostly complete list of all taxa described up until 2018 can be found in Ross 2018; its supplement Ross 2019b covers most of 2019.

Umenocoleidae is an extinct family of dictyopteran insects known from the Cretaceous. They are considered to be closely related to the Alienopteridae. They were originally considered to be beetles due to their beetle-like morphology, with sclerotised elytra-like forewings. This was probably an adaptation for living under bark and in other tight spaces.

<span class="mw-page-title-main">Babinskaiidae</span> Extinct family of insects

Babinskaiidae is an extinct family of neuropterans known from the Cretaceous period. They are part of the superfamily Myrmeleontoidea. Their distinguishing characters include: "long filiform antennae, narrowly elongated wings, with features such as trichosors, and presectorial cross veins present in both wings, and absence of forewing oblique vein". They are considered transitional between Nymphidae and more derived myrmeleontodoids, such as antlions.

<span class="mw-page-title-main">Zhangsolvidae</span> Extinct family of flies

Zhangsolvidae is an extinct family of brachyceran flies known from the Cretaceous period. Members of the family possess a long proboscis, varying in length between 1.3 and 7 mm depending on the species, and were probably nectarivores. A specimen has been found with preserved Bennettitales pollen, suggesting that they acted as pollinators for extinct gymnosperms. They are considered to be members of the Stratiomyomorpha.

<span class="mw-page-title-main">Elcanidae</span> Extinct family of cricket-like animals

Elcanidae are an extinct family of Mesozoic and early Cenozoic orthopterans. Members of the family are distinguished by the presence of spurs on the distal part of the metatibia, unique among orthopterans, these have been suggested to have been used for controlling gliding, swimming aids, or for jumping on water. The group combines characteristics from both major groups of orthopterans, with long antennae and nymphal morphology similar to Ensifera, but with wing venation and adult morphology more similar to Caelifera. Elcanidae is part of Elcanoidea, which is thought to have diverged from living orthopterans by the beginning of the Permian, around 300 million years ago. The family also includes Permelcanidae, known from the Early-Late Permian. The relationship of Elcanoidea to Ensifera and Caelifera is currently unresolved. Elcanids are known from the Late Triassic to Paleocene of Eurasia, North and South America. Some members of the group exhibited aposematic coloration.

This paleoentomology list records new fossil insect taxa that were to be described during the year 2021, as well as notes other significant paleoentomology discoveries and events which occurred during that year.

This paleoentomology list records new fossil insect taxa that are to be described during the year 2022, as well as notes other significant paleoentomology discoveries and events which occurred during that year.

This list of 2023 in paleoentomology records new fossil insect taxa that are to be described during the year, as well as documents significant paleoentomology discoveries and events which occurred during that year.

References

  1. Mey, Wolfram; Wichard, Wilfried; Ross, Emma; Ross, Andrew (January 2018). "On the systematic position of a highly derived amphiesmenopteran insect from Burmese amber (Insecta, Amphiesmenoptera)". Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 107 (2–3): 249–254. doi:10.1017/S1755691017000330. ISSN   1755-6910. S2CID   88661747.
  2. 1 2 3 Mey, Wolfram; Wichard, Wilfried; Müller, Patrick; Wang, Bo (2017-03-24). "The blueprint of the Amphiesmenoptera Tarachoptera, a new order of insects from Burmese amber (Insecta, Amphiesmenoptera)". Fossil Record. 20 (2): 129–145. doi: 10.5194/fr-20-129-2017 . ISSN   2193-0074.
  3. Mey, W.; Wichard, W.; Müller, P.; Ross, E.; Ross, A. (October 2018). "New taxa of Tarachoptera from Burmese amber (Insecta, Amphiesmenoptera)". Cretaceous Research. 90: 154–162. doi:10.1016/j.cretres.2018.04.006. S2CID   133848887.
  4. Mey, Wolfram; Wichard, Wilfried; Müller, Patrick; Wang, Bo (2020-06-23). "Descriptions of two new species of Tarachoptera from Burmese amber (Insecta, Amphiesmenoptera: Tarachoptera)". Contributions to Entomology = Beiträge zur Entomologie. 70: 181–188 Seiten. doi: 10.21248/CONTRIB.ENTOMOL.70.1.181-188 .
  5. Wang, Jiajia; Zhang, Weiting; Engel, Michael S.; Sheng, Xianyong; Shih, Chungkun; Ren, Dong (September 2022). "Early evolution of wing scales prior to the rise of moths and butterflies". Current Biology. 32 (17): 3808–3814.e2. doi: 10.1016/j.cub.2022.06.086 . PMID   35998638. S2CID   251727701.