Tortoises are reptiles of the family Testudinidae of the order Testudines. They are particularly distinguished from other turtles by being exclusively land-dwelling, while many other turtle species are at least partly aquatic. Like other turtles, tortoises have a shell to protect from predation and other threats. The shell in tortoises is generally hard, and like other members of the suborder Cryptodira, they retract their necks and heads directly backwards into the shell to protect them.
Emydidae is a family of testudines (turtles) that includes close to 50 species in 10 genera. Members of this family are commonly called terrapins, pond turtles, or marsh turtles. Several species of Asian box turtles were formerly classified in the family; however, revised taxonomy has separated them to a different family (Geoemydidae). As currently defined, the Emydidae are entirely a Western Hemisphere family, with the exception of two species of pond turtle.
The mata mata, mata-mata, or matamata is a freshwater turtle species found in South America, primarily in the Amazon and Orinoco basins. It is one of two extant species in the genus Chelus, the other being Chelus orinocensis.
The marginated tortoise is a species of tortoise in the family Testudinidae. The species is endemic to Greece, Italy, and the Balkans in Southern Europe. It is the largest European tortoise. The marginated tortoise is herbivorous, and hibernates for the winter.
Hermann's tortoise is a species of tortoise. Two subspecies are known: the western Hermann's tortoise and the eastern Hermann's tortoise. Sometimes mentioned as a subspecies, T. h. peleponnesica is not yet confirmed to be genetically different from T. h. boettgeri.
The red-footed tortoise is a species of tortoise from northern South America. These medium-sized tortoises generally average 30 cm (12 in) as adults, but can reach over 40 cm (16 in). They have dark-colored, loaf-shaped carapaces with a lighter patch in the middle of each scute, and dark limbs with brightly colored scales that range from pale yellow to dark red. Recognized differences are seen between red-footed tortoises from different regions. They are closely related to the yellow-footed tortoise from the Amazon Basin. They are popularly kept as pets, and over-collection has caused them to be vulnerable to extinction.The species name has often been misspelled as carbonaria, an error introduced in the 1980s when Chelonoidis was elevated to genus and mistakenly treated as feminine, an error recognized and fixed in 2017.
The Asian forest tortoise, also known commonly as the Asian brown tortoise, is a species of tortoise in the family Testudinidae. The species is endemic to Southeast Asia. It is believed to be among the most primitive of living tortoises, based on molecular and morphological studies.
Testudo, the Mediterranean tortoises, are a genus of tortoises found in North Africa, Western Asia, and Europe. Several species are under threat in the wild, mainly from habitat destruction.
Geochelone is a genus of tortoises.
Pelusios is a genus of African side-necked turtles. With 17 described species, it is one of the most diverse genera of the turtle order (Testudines).
The Tunisian tortoise or Nabeul tortoise is a subspecies of Greek tortoises. It was originally described as a new species in 1990, and even placed in a distinct genus. The spur-thighed or "Greek" tortoises are usually collectively referred to as Testudo graeca, but this covers a wide variety of subspecies that have very different ecological and morphological characteristics and appear to comprise at least three phylogenetic lineages. As its name implies, it is found in Tunisia and nearby Algeria.
The Galápagos tortoise complex or Galápagos giant tortoise complex is a species complex of 15 very large tortoise species in the genus Chelonoidis. They are the largest living species of tortoise, with some modern Galápagos tortoises weighing up to 417 kg (919 lb). With lifespans in the wild of over 100 years, they are one of the longest-lived vertebrates. Captive Galapagos tortoises can live up to 177 years. For example, a captive individual, Harriet, lived for at least 175 years. Spanish explorers, who discovered the islands in the 16th century, named them after the Spanish galápago, meaning "tortoise".
Kleinmann's tortoise, also called commonly the Egyptian tortoise, Leith's tortoise, and the Negev tortoise, is a critically endangered species of neck-hiding tortoise in the family Testudinidae. The species is native to Egypt, Libya, and Israel. The species was once more widespread, but its numbers are now dwindling. The species is nearly extinct in Egypt, and complete extinction in the wild is a looming threat unless more actions are taken to protect this species.
The yellow-footed tortoise, also known as the Brazilian giant tortoise, commonly referred to as the Brazilian giant turtle, or more commonly, the big turtle, is a species of tortoise in the family Testudinidae and is closely related to the red-footed tortoise. It is found in the Amazon Basin of South America. The species name has often been misspelled as denticulata, an error introduced in the 1980s when Chelonoidis was elevated to genus and mistakenly treated as feminine, an error recognized and fixed in 2017.
Hesperotestudo is an extinct genus of tortoise native to North and Central America from the Oligocene to the Late Pleistocene. Species of Hesperotesudo varied widely in size, with the largest species reaching sizes comparable to extant giant tortoises, with carapaces over a metre in length. Historically considered a subgenus of Geochelone, it is now considered to be distantly related to that genus. Its closest relatives are the extant Gopherus and the extinct Stylemys. The ancestor of the three genera arrived in North America during the Early Eocene. The bodies of Hesperotesudo species were extensively covered with large dermal ossicles, which in life were covered in keratin. It has been suggested that species of Hesperotestudo were relatively tolerant of cold weather. Hesperotestudo became extinct at the end of the Pleistocene and beginning of the Holocene co-incident with the arrival of the first humans in North America, and sites have been found where Hesperotestudo were butchered.
The Astaracian age is a period of geologic time, equivalent with the Middle Miocene and used more specifically with European Land Mammal Ages. It precedes the Vallesian age and follows the Orleanian age. The Astaracian overlaps the Langhian and Serravallian ages.
Chelodina canni, also known commonly as Cann's snake-necked turtle, is a species of turtle in the family Chelidae. The species is endemic to Australia, where it is found in the northern and northeastern parts of the continent. It has a narrow zone of hybridization with its related species the eastern snake-necked turtle, C. longicollis. For many years C. canni was assumed to be the same species as C. novaeguineae from New Guinea. However, in 2002 it was shown that these two species differ both morphologically and genetically, and therefore C. canni was separated and described as a unique species.
Chelonoidis is a genus of turtles in the tortoise family erected by Leopold Fitzinger in 1835. They are found in South America and the Galápagos Islands, and formerly had a wide distribution in the West Indies.
Megalochelys is an extinct genus of cryptodiran tortoises that lived from the Miocene to Pleistocene. They are noted for their giant size, which is among the largest of any known testudine, with a maximum carapace length over 2 m (6.5 ft) in M. atlas. During the dry glacial periods it ranged from western India and Pakistan to as far east as Sulawesi and Timor in Indonesia, though the island specimens likely represent distinct species.
Titanochelon is an extinct genus of giant tortoises known from the Early Miocene to the beginning of the Pleistocene in Europe, extending from the Iberian Peninsula to Anatolia. Some members of the genus were larger than extant giant tortoises, with a shell length of up to 2 metres.
