Thalattosauroidea

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Thalattosauroidea
Temporal range: Middle To Late Triassic, 247.2–215  Ma
Thalattosauridae.jpg
Life restoration of Nectosaurus halinus (right) and Thalattosaurus alexandrae (left)
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Thalattosauria
Superfamily: Thalattosauroidea
Merriam, 1904
Subtaxa
Synonyms

Thalattosauroidea is a superfamily of thalattosaurs, a Triassic group of marine reptiles. It was named in 1904 by paleontologist John Campbell Merriam to include the genus Thalattosaurus from California. Thalattosauroids are one of two groups of Thalattosauria, the other being Askeptosauroidea. Thalattosauroids make up the "traditional" thalattosaurs with large downturned snouts, short necks, and long, paddle-like tails.

Classification

Thalattosauria includes North American genera such as Thalattosaurus and Nectosaurus as well as recently described Chinese forms such as Xinpusaurus . A 1999 study of thalattosaurs, which established much of the currently accepted phylogeny of the group, referred to Thalattosauroidea as Thalattosauria, while calling the larger group Thalattosauriformes. More recent phylogenetic studies have come upon the same conclusions, but refer to the group as Thalattosauroidea in order to contrast it with another superfamily of thalattosaurs, Askeptosauroidea. [1] [2] Below is a cladogram from Wu et al. (2009) showing the phylogenetic relationships of Thalattosauroidea: [3]

Thalattosauria

Related Research Articles

<i>Dinocephalosaurus</i> Extinct genus of reptiles

Dinocephalosaurus is a genus of long necked, aquatic protorosaur that inhabited the Triassic seas of China. The genus contains the type and only known species, D. orientalis, which was named by Chun Li in 2003. Unlike other long-necked protorosaurs, Dinocephalosaurus convergently evolved a long neck not through elongation of individual neck vertebrae, but through the addition of neck vertebrae that each had a moderate length. As indicated by phylogenetic analyses, it belonged in a separate lineage that also included at least its closest relative Pectodens, which was named the Dinocephalosauridae in 2021. Like tanystropheids, however, Dinocephalosaurus probably used its long neck to hunt, utilizing the fang-like teeth of its jaws to ensnare prey; proposals that it employed suction feeding have not been universally accepted. It was probably a marine animal by necessity, as suggested by the poorly-ossified and paddle-like limbs which would have prevented it from going ashore.

<i>Nectosaurus</i> Extinct genus of reptiles

Nectosaurus is a genus of thalattosaur which lived during the Late Triassic in what is now California. The type and only known species, Nectosaurus halius, was found in the Hosselkus Limestone and described by John C. Merriam in 1905, making it one of the first thalattosaurians known.

<i>Askeptosaurus</i> Extinct genus of reptiles

Askeptosaurus is an extinct genus of askeptosauroid, a marine reptile from the extinct order Thalattosauria. Askeptosaurus is known from several well-preserved fossils found in Middle Triassic marine strata in what is now Italy and Switzerland.

<span class="mw-page-title-main">Thalattosauria</span> Extinct group of marine reptiles

Thalattosauria is an extinct order of marine reptiles that lived in the Middle to Late Triassic. Thalattosaurs were diverse in size and shape, and are divided into two superfamilies: Askeptosauroidea and Thalattosauroidea. Askeptosauroids were endemic to the Tethys Ocean, their fossils have been found in Europe and China, and they were likely semiaquatic fish eaters with straight snouts and decent terrestrial abilities. Thalattosauroids were more specialized for aquatic life and most had unusual downturned snouts and crushing dentition. Thalattosauroids lived along the coasts of both Panthalassa and the Tethys Ocean, and were most diverse in China and western North America. The largest species of thalattosaurs grew to over 4 meters (13 feet) in length, including a long, flattened tail utilized in underwater propulsion. Although thalattosaurs bore a superficial resemblance to lizards, their exact relationships are unresolved. They are widely accepted as diapsids, but experts have variously placed them on the reptile family tree among Lepidosauromorpha, Archosauromorpha, ichthyosaurs, and/or other marine reptiles.

<i>Miodentosaurus</i> Extinct genus of reptiles

Miodentosaurus is a genus of thalattosaurian from the Late Triassic of China. It is one of several thalattosaurs found in the Xiaowa Formation, also known as the Wayao Member of the Falang Formation. The genus name "Miodentosaurus" translates to "Few toothed-lizard" while the species name "brevis" means "short", in reference to its short snout.

<i>Thalattosaurus</i> Extinct genus of reptiles

Thalattosaurus meaning "sea lizard," from the Attic Greek thalatta (θάλαττα), "sea," and sauros (σαῦρος), "lizard," is an extinct genus of marine reptile in the family Thalattosauroidea. They were aquatic diapsids that are known exclusively from the Triassic period. It was a 2–3 metres (6.6–9.8 ft) long shellfish-eating reptile with paddle-like limbs and a down-turned rostrum occurring in the Lower and Middle Triassic Sulphur Mountain Formation of British Columbia as well as the Upper Triassic Hosselkus Limestone of California. It has gained notoriety as a result of studies on general diapsid phylogeny.

<i>Helveticosaurus</i> Extinct genus of reptiles

Helveticosaurus is an extinct genus of diapsid marine reptile known from the Middle Triassic of southern Switzerland. It contains a single species, Helveticosaurus zollingeri, known from the nearly complete holotype T 4352 collected at Cava Tre Fontane of Monte San Giorgio, an area well known for its rich record of marine life during the Middle Triassic.

The Guanling Formation is a Middle Triassic geologic formation in southwestern China.

<i>Sinosaurosphargis</i> Extinct genus of reptiles

Sinosaurosphargis is an extinct genus of basal marine saurosphargid reptile known from the Middle Triassic Guanling Formation of Yunnan and Guizhou Provinces, southwestern China. It contains a single species, Sinosaurosphargis yunguiensis.

<span class="mw-page-title-main">Askeptosauroidea</span> Extinct superfamily of reptiles

Askeptosauroidea is a superfamily of thalattosaurs, a Triassic group of marine reptiles. Askeptosauroidea is one of two major subgroups of Thalattosauria, the other being Thalattosauroidea. It includes the family Askeptosauridae and a more basal form called Endennasaurus.

<span class="mw-page-title-main">Askeptosauridae</span> Extinct family of reptiles

Askeptosauridae is a family of thalattosaurs within the superfamily Askeptosauroidea. Fossils have been found from Italy, Switzerland, and China. Askeptosaurids are distinguished from other thalattosaurs by their long necks and narrow skulls.

<i>Anshunsaurus</i> Extinct genus of reptiles

Anshunsaurus is a genus of thalattosaurs within the family Askeptosauridae. Fossils have been found from Middle Triassic deposits in Guizhou, China. Three species are known: the type species A. huangguoshuensis, the slightly older species A. wushaensis, and the species A. huangnihensis.

<i>Wumengosaurus</i> Extinct genus of reptiles

Wumengosaurus is an extinct aquatic reptile from the Middle Triassic Guanling Formation of Guizhou, southwestern China. It was originally described as a basal eosauropterygian and usually is recovered as such by phylogenetic analyses, although one phylogeny has placed it as the sister taxon to Ichthyosauromorpha while refraining from a formal re-positioning. It was a relatively small reptile, measuring 95.5–130.5 cm (3.13–4.28 ft) in total body length.

Hanosaurus is an extinct genus of marine reptiles that existed during the Triassic period in what is now China. The type species is Hanosaurus hupehensis. It was a small animal, with specimens measuring 79.4 cm (31.3 in) long in total body length, which likely fed on soft-bodied prey.

<i>Concavispina</i> Extinct genus of reptiles

Concavispina is an extinct genus of thalattosaur reptile from the early Late Triassic Xiaowa Formation of Guangling, Guizhou, southern China. It contains a single species, Concavispina biseridens. It is known only from the holotype ZMNH M8804, a nearly complete 364 cm long skeleton. Concavispina can be differentiated from other thalattosaurs by possessing two rows of blunt teeth on the anterior part of the maxilla and a V-shaped notch on the dorsal margin of each neural spine in the dorsal (back) vertebrae. Both its generic and specific names refer to these autapomorphies, as Concavispina means "concave spine" and biseridens means "two rows of teeth". It is thought to be most closely related to Xinpusaurus, as both taxa share three derived characters: a maxilla that is curved upward at its anterior end, a humerus that is wider near the shoulder than near the elbow, and the presence of less than five cervicals.

<i>Xinpusaurus</i> Extinct genus of reptiles

Xinpusaurus is an extinct genus of thalattosaur from the Late Triassic of Guanling in Guizhou, China. Several species have been named since 2000: the type species X. suni along with the species X. bamaolinensis and X. kohi. A 2013 study proposed that all three species are synonymous with each other, in which case X. suni would be the only valid species, although a 2014 study argued that X. kohi was also valid. A fourth species, X. xingyiensis, was described in 2016.

<i>Blezingeria</i> Extinct genus of reptiles

Blezingeria is an extinct genus of marine reptile from the Middle Triassic of Germany. The type and only species Blezingeria ichthyospondyla was named by German paleontologist Eberhard Fraas in 1896. It is known from many isolated bones that come from a deposit in southwestern Germany called the Upper Muschelkalk, which dates back to the Ladinian stage. The relationships of Blezingeria are uncertain. Fraas identified it as a nothosaur, but it has also been classified as a cymbospondylid ichthyosaur, and most recently a thalattosaur. Many thalattosaur fossils have been found in a slightly olderLate Ladinian-age rock unit in Monte San Giorgio, Switzerland, so if Blezingeria is a thalattosaur, it may represent an early stage in an evolutionary radiation of the group across the Tethys, an ocean that covered much of what is now Europe and southern China during the Triassic. However, since Blezingeria is known from very incomplete material, its classification as a thalattosaur remains uncertain.

<span class="mw-page-title-main">Saurosphargidae</span> Extinct family of reptiles

Saurosphargidae is an extinct family of marine reptiles known from the Early Triassic and early Middle Triassic of Europe and China.

The Xiaowa Formation is a Carnian-age geological formation found in southern China. It is a sequence of limestone and marls from the Carnian stage of the Triassic. Its lower section was previously known as the Wayao Formation or Wayao Member of the Falang Formation. In 2002, the Wayao Member was renamed and raised to the Xiaowa Formation to prevent confusion with an Eocene unit of the same name. Crinoids and marine reptiles are abundant in the Xiaowa Formation, forming a lagerstätte known as the Guanling biota. Ammonoids and conodonts found in the formation constrain its age to the early Carnian. Reptiles of the Guanling biota include ichthyosaurs, thalattosaurs, placodonts, and Odontochelys. Sedimentary events within this formation have been tied to the Carnian Pluvial Event.

The Zhuganpo Formation is a Triassic geologic unit found in southern China. It has historically been known as the Zhuganpo Member of the Falang Formation. A diverse fossil assemblage known as the Xingyi biota or Xingyi Fauna can be found in the upper part of the Zhuganpo Formation. Fossils of the Xingyi biota include articulated skeletons of marine reptiles, abundant fish, and a plentiful assortment of invertebrates indicating a Ladinian to Carnian age for the sediments of the formation.

References

  1. Rieppel, O.; Liu, J.; Bucher, H. (2000). "The first record of a thalattosaur reptile from the Late Triassic of southern China (Guizhou Province, PR China)". Journal of Vertebrate Paleontology. 20 (3): 507–514. doi:10.1671/0272-4634(2000)020[0507:TFROAT]2.0.CO;2. S2CID   140706921.
  2. Liu, J.; Rieppel, O. (2005). "Restudy of Anshunsaurus huangguoshuensis (Reptilia: Thalattosauria) from the Middle Triassic of Guizhou, China" (PDF). American Museum Novitates (3488): 1–34. doi:10.1206/0003-0082(2005)488[0001:roahrt]2.0.co;2. ISSN   0003-0082. S2CID   55642315.
  3. Wu, X.-C.; Cheng, Y.-N.; Sato, T.; Shan, H.-Y. (2009). "Miodentosaurus brevis Cheng et al., 2007 (Diapsida: Thalattosauria): its postcranial skeleton and phylogenetic relationships" (PDF). Vertebrata PalAsiatica. 47 (1): 1–20.