Treptichnus Temporal range: | |
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Burrows viewed from the side | |
Burrows viewed obliquely | |
Trace fossil classification | |
Ichnogenus: | † Treptichnus Miller, 1889 |
Ichnospecies | |
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Synonyms [2] [3] | |
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Treptichnus (formerly named Phycodes, Manykodes by J. Dzik, [2] and also known as Trichophycus [3] ) is the preserved burrow of an animal. As such, it is regarded as the earliest widespread complex trace fossil. Its earliest appearance, around 542 million years ago (mya), [4] which was contemporaneous with the last of the Ediacaran biota, is used to help define the dividing line, considered geologically at 541 mya, between the Ediacaran and Cambrian periods. [5] [6] It is last seen in the fossil record during the Cenomanian (99.7 mya). [1]
Treptichnus pedum has a fairly complicated and distinctive burrow pattern: along a central, sometimes sinuous or looping burrow it made successive probes upward through the sediment in search of nutrients, generating a trace pattern reminiscent of a fan or twisted rope. [7] The organism that left such traces is considered more complex than earlier Ediacaran biota; and these trace fossils, which occur worldwide, are usually found in strata above those. [8]
Since only its burrows have been found, it is presumed that the Treptichnus animal lacked any hard anatomical features, such as shells or bones. Its morphology and relationship to modern animals is therefore unknown, and some dispute even its inclusion within the animal kingdom. [9] Despite this lack of fossil evidence, surface trails produced by modern priapulid worms exhibit the same probing morphology as Treptichnus, suggesting a close anatomical relationship between the trace-maker and modern priapulids. [10]
The name Treptichnus pedum means "turned-trail (Greek) of feet (Latin)".
The Ediacaran is a geological period of the Neoproterozoic Era that spans 96 million years from the end of the Cryogenian Period at 635 Mya to the beginning of the Cambrian Period at 538.8 Mya. It is the last period of the Proterozoic Eon as well as the last of the so-called "Precambrian supereon", before the beginning of the subsequent Cambrian Period marks the start of the Phanerozoic Eon, where recognizable fossil evidence of life becomes common.
The Neoproterozoic Era is the unit of geologic time from 1 billion to 538.8 million years ago.
A trace fossil, also known as an ichnofossil, is a fossil record of biological activity by lifeforms but not the preserved remains of the organism itself. Trace fossils contrast with body fossils, which are the fossilized remains of parts of organisms' bodies, usually altered by later chemical activity or by mineralization. The study of such trace fossils is ichnology - the work of ichnologists.
Vendobionts or Vendozoans (Vendobionta) are a proposed very high-level, extinct clade of benthic organisms that made up of the majority of the organisms that were part of the Ediacaran biota. It is a hypothetical group and at the same time, it would be the oldest of the animals that populated the Earth about 580 million years ago, in the Ediacaran period. They became extinct shortly after the so-called Cambrian explosion, with the introduction of fauna forming groups more recognizably related to modern animals. It is very likely that the whole Ediacaran biota is not a monophyletic clade and not every genus placed in its subtaxa is an animal.
Fortune Head is a headland located about 1.6 km (0.99 mi) from the town of Fortune on the Burin Peninsula, southeastern Newfoundland.
Yorgia waggoneri is a discoid Ediacaran organism. It has a low, segmented body consisting of a short wide "head", no appendages, and a long body region, reaching a maximum length of 25 cm (9.8 in). It is classified within the extinct animal phylum Proarticulata.
The Ediacaranbiota is a taxonomic period classification that consists of all life forms that were present on Earth during the Ediacaran Period. These were enigmatic tubular and frond-shaped, mostly sessile, organisms. Trace fossils of these organisms have been found worldwide, and represent the earliest known complex multicellular organisms. The term "Ediacara biota" has received criticism from some scientists due to its alleged inconsistency, arbitrary exclusion of certain fossils, and inability to be precisely defined.
The "Cambrian substrate revolution" or "Agronomic revolution", evidenced in trace fossils, is a sudden diversification of animal burrowing during the early Cambrian period.
The small shelly fauna, small shelly fossils (SSF), or early skeletal fossils (ESF) are mineralized fossils, many only a few millimetres long, with a nearly continuous record from the latest stages of the Ediacaran to the end of the Early Cambrian Period. They are very diverse, and there is no formal definition of "small shelly fauna" or "small shelly fossils". Almost all are from earlier rocks than more familiar fossils such as trilobites. Since most SSFs were preserved by being covered quickly with phosphate and this method of preservation is mainly limited to the late Ediacaran and early Cambrian periods, the animals that made them may actually have arisen earlier and persisted after this time span.
The Cambrian explosion is an interval of time approximately 538.8 million years ago in the Cambrian period of the early Paleozoic when a sudden radiation of complex life occurred, and practically all major animal phyla started appearing in the fossil record. It lasted for about 13 to 25 million years and resulted in the divergence of most modern metazoan phyla. The event was accompanied by major diversification in other groups of organisms as well.
The Stratigraphy of the Cambrian period currently has several schemes used for ordering geologic formations from the period. The International Commission on Stratigraphy−ICS scheme has set a stratotype section for the base of the Cambrian, dated quite accurately to 538.8 ± 0.2 million years ago. Russian and Chinese scientists have developed a different scheme.
Martin David Brasier FGS, FLS was an English palaeobiologist and astrobiologist known for his conceptual analysis of microfossils and evolution in the Precambrian and Cambrian.
The Terreneuvian is the lowermost and oldest series of the Cambrian geological system. Its base is defined by the first appearance datum of the trace fossil Treptichnus pedum around 538.8 million years ago. Its top is defined as the first appearance of trilobites in the stratigraphic record around 521 million years ago. This series' name was formally accepted by the International Commission on Stratigraphy in 2007.
The Fortunian age marks the beginning of the Phanerozoic Eon, the Paleozoic Era, and the Cambrian Period. It is the first of the two stages of the Terreneuvian series. Its base is defined as the first appearance of the trace fossil Treptichnus pedum 538.8 million years ago. The top of the Fortunian which is the base of the Stage 2 of the Cambrian has not been formally defined yet, but will correspond to the appearance of an Archeocyatha species or "Small shelly fossils" approximately 529 million years ago.
The Chapel Island Formation is a sedimentary formation from the Burin Peninsula, Newfoundland, Canada. It is a succession of siliciclastic deposits, over 1,000 metres (3,300 ft) thick, that were deposited during the latest Ediacaran and earliest Cambrian.
The Nama Group is a 125,000 square kilometres (48,000 sq mi) megaregional Vendian to Cambrian group of stratigraphic sequences deposited in the Nama foreland basin in central and southern Namibia. The Nama Basin is a peripheral foreland basin, and the Nama Group was deposited in two early basins, the Zaris and Witputs, to the north, while the South African Vanrhynsdorp Group was deposited in the southern third. The Nama Group is made of fluvial and shallow-water marine sediments, both siliciclastic and carbonate. La Tinta Group in Argentina is considered equivalent to Nama Group.
Jerzy Dzik is a Polish paleontologist.
The Ediacaran fossil Hallidaya, a close relative of Skinnera lived in Belomorian of the Late Ediacaran period prior to the Cambrian explosion and thrived in the marine strata on the ocean floor of what is now considered Australia. These fossils were disk-shaped organisms that were slightly dome shaped with tri-radial symmetry. These Ediacaran organisms thrived by living in low-energy inner shelf, in the wave- and current-agitated shoreface, and in the high-energy distributary systems.
Matgrounds are strong surface layers of seabed-hardening bacterial fauna preserved in the Proterozoic and lower Cambrian. Wrinkled matgrounds are informally named "elephant skin" because of its wrinkled surface in the fossil record. Matgrounds supported themselves until early burrowing worms were ubiquitous enough to unharden them. Burrowing animals broke down the hardy mats to further penetrate the underlying sediment for protection and feeding. Once matgrounds disappeared, exceptional preservation of lagerstätten such as the Burgess Shale or Ediacara Hills also did so too. Trace fossils such as Treptichnus are evidence for soft-bodied burrowers more anatomically complex than the Ediacaran biota that also caused the matgrounds disappearance.