| Xiaoheiqingella Temporal range: | |
|---|---|
 | |
| Reconstruction | |
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Priapulida |
| Family: | † Xiaoheiqingidae |
| Genus: | † Xiaoheiqingella |
| Species: | †X. peculiaris |
| Binomial name | |
| †Xiaoheiqingella peculiaris Hu in Chen et al. 2002 | |
Xiaoheiqingella is a taxon of priapulid known from the Chengjiang biota; synonymous with Yunnanpriapulus, [1] and thought to belong to the priapulid crown group. [2]
Priapulida, sometimes referred to as penis worms, is a phylum of unsegmented marine worms. The name of the phylum relates to the Greek god of fertility, because their general shape and their extensible spiny introvert (eversible) proboscis may resemble the shape of a human penis. They live in the mud and in comparatively shallow waters up to 90 metres (300 ft) deep. Some species show a remarkable tolerance for hydrogen sulfide, anoxia and low salinity. Halicryptus spinulosus appears to prefer brackish shallow waters. They can be quite abundant in some areas. In an Alaskan bay as many as 85 adult individuals of Priapulus caudatus per square meter has been recorded, while the density of its larvae can be as high as 58,000 per square meter.
In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. It is thus a way of defining a clade, a group consisting of a species and all its extant or extinct descendants. For example, Neornithes (birds) can be defined as a crown group, which includes the most recent common ancestor of all modern birds, and all of its extant or extinct descendants.
Ottoia is a stem-group archaeopriapulid worm known from Cambrian fossils. Although priapulid-like worms from various Cambrian deposits are often referred to Ottoia on spurious grounds, the only clear Ottoia macrofossils come from the Burgess Shale of British Columbia, which was deposited 508 million years ago. Microfossils extend the record of Ottoia throughout the Western Canada Sedimentary Basin, from the mid- to late- Cambrian. A few fossil finds are also known from China.
Dinocaridida is a proposed fossil taxon of basal arthropods that flourished in the Cambrian period with occasional Ordovician and Devonian records. Characterized by a pair of frontal appendages and series of body flaps, the name of Dinocaridids refers to the suggested role of some of these members as the largest marine predators of their time. Dinocaridids are occasionally referred to as the 'AOPK group' by some literatures, as the group compose of Radiodonta, Opabiniidae, and the "gilled lobopodians" Pambdelurion and Kerygmachelidae. It is most likely paraphyletic, with Kerygmachelidae and Pambdelurion more basal than the clade compose of Opabiniidae, Radiodonta and other arthropods.
Treptichnus is the preserved burrow of an animal. As such, it is regarded as the earliest widespread complex trace fossil. Its earliest appearance, around 542 mya, which was contemporaneous with the last of the Ediacaran biota, is used to help define the dividing line, considered geologically at 541 mya, between the Ediacaran and Cambrian periods. It is last seen in the fossil record during the Cenomanian.
Fieldia is a genus of worms known from the Cambrian Burgess Shale, and assigned to the priapulids.
Facivermis is a genus of sessile lobopodian from the Lower Cambrian Maotianshan shales of China
Jianshanopodia is a monotypic genus of Cambrian lobopodian, discovered from Maotianshan Shales of Yunnan, China.
Ancalagon minor is an extinct priapulid worm known from the Cambrian Burgess Shale.
Archaeopriapulida is a group of priapulid worms known from Cambrian lagerstätte. The group is closely related to, and very similar to, the modern Priapulids. It is unclear whether it is mono- or polyphyletic. Despite a remarkable morphological similarity to their modern cousins, they fall outside of the priapulid crown group, which is not unambiguously represented in the fossil record until the Carboniferous. In addition to well-preserved body fossils, remains of several archaeopriapulid taxa are known to have been preserved primarily as organic microfossils, such as isolated scalids and pharyngeal teeth. They are probably closely related or paraphyletic to the palaeoscolecids; the relationship between these basal worms is somewhat unresolved.
The palaeoscolecids are a group of extinct ecdysozoan worms resembling armoured priapulids. They are known from the Lower Cambrian to the late Silurian; they are mainly found as disarticulated sclerites, but are also preserved in many of the Cambrian lagerstätten. They take their name from the typifying genus Palaeoscolex. Other genera include Cricocosmia from the Lower Cambrian Chengjiang biota. Their taxonomic affinities within Ecdysozoa have been the subject of debate.
Cambrorhytium is an enigmatic fossil genus known from the Latham Shale (California), and the Chengjiang (China) and Burgess Shale lagerstätte. 350 specimens of Cambrorhytium are known from the Greater Phyllopod bed, where they comprise 0.7% of the community.
Selkirkia is a genus of predatory, tubicolous priapulid worms known from the Middle Cambrian Burgess Shale, Ogygopsis Shale and Puncoviscana Formation. 142 specimens of Selkirkia are known from the Greater Phyllopod bed, where they comprise 0.27% of the community. In the Burgess Shale, 20% of the tapering, organic-walled tubes are preserved with the worm inside them, whereas the other 80% are empty. Whilst alive, the tubes were probably vertical, whereas trilobite-occupied tubes are horizontal.
Omnidens amplus, meaning "large all-tooth", is an extinct species of large Cambrian animal known only from a series of large mouth apparatus, originally mistaken as the mouthparts of anomalocaridids. When first named, it was interpreted as a giant priapulid, but is now considered a panarthropod. Its mouth apparatus closely resembles that of the smaller gilled lobopodian Pambdelurion, indicating it is likely to have been a close relative of that species, with which it may be synonymous. With a maximum estimated body length of 1.5 metres (4.9 ft), Omnidens is suggested to have been the largest known free-living Cambrian organism. Omnidens fossils are found in the Maotianshan Shales.
Small carbonaceous fossils (SCFs) are sub-millimetric organic remains of organisms preserved in sedimentary strata.
Acosmia is an extinct genus of marine worm from the Cambrian aged Chengjiang biota of Yunnan, China. It is represented by a single rare species, Acosmia maotiania. It was likely a burrowing animal that fed by deposit feeding. While originally suggested to be a priapulid, a 2020 study proposed to be a stem-group ecdysozoan.
Sicyophorus is a genus of archaeopriapulid known from the Chengjiang biota; synonymous with Protopriapulites haikouensis.
Paraselkirkia is a genus of archaeopriapulid known from the Chengjiang biota, resembling Selkirkia.
Eximipriapulus is a genus of priapulid-like organisms, perhaps belonging to the crown group, known from the Chengjiang biota.
Corynetis is a genus of archaeopriapulid known from the Chengjiang biota, and a senior synonym of Anningvermis multispinosus.
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