Andropogoneae | |
---|---|
Andropogon scoparius | |
Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Angiosperms |
Clade: | Monocots |
Clade: | Commelinids |
Order: | Poales |
Family: | Poaceae |
Clade: | PACMAD clade |
Subfamily: | Panicoideae |
Supertribe: | Andropogonodae |
Tribe: | Andropogoneae Dumort. (1824) |
Subtribes | |
12 subtribes, see text | |
Synonyms [1] | |
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The Andropogoneae, sometimes called the sorghum tribe, are a large tribe of grasses (family Poaceae) with roughly 1,200 species in 90 genera, mainly distributed in tropical and subtropical areas. They include such important crops as maize (corn), sugarcane, and sorghum. [2] All species in this tribe use C4 carbon fixation, which makes them competitive under warm, high-light conditions. [3]
Andropogoneae is classified in supertribe Andropogonodae, together with its sister group Arundinelleae. Subdivisions include 12 subtribes, but the position of several genera within them is still unresolved ( incertae sedis ). Hybridisation was probably important in the evolution of the Andropogoneae, and the tribe's systematics are still not completely resolved. [3] From the morphological point of view, the merging of the former subtribe 'Dimeriinae' (having solitary spikelets borne in 'robust' raceme rachis ) into the subtribe Ischaeminae (having paired spikelets borne in 'fragile' raceme rachis) is doubtful. Affinities within the tribe are complex and still under investigation. [4] Mr. Shahid Nawaz, a grass specialist at the Blatter Herbarium, is working on the phylogenomics of the tribe from India.[ citation needed ]
According to preliminary phytogeographic research on the tribe by MS Kiran Raj, Peninsular India, home to 54 genera (incl. 9 endemic genera, viz., Bhidea, Lophopogon, Glyphochloa, Pogonachne, Trilobachne, Pseudodichanthium, Triplopogon, Nanooravia, and Manisuris ) and roughly 500 species, is thought to be the primary or secondary centre of diversity for Andropogonoid grasses. About 40% of the representative taxa are exclusively endemic to peninsular India, with the highest species diversity and endemicity found in genera like Arthraxon, Chrysopogon , Cymbopogon , Dichanthium , Dimeria , Heteropogon, Ischaemum, Ophiorus, Sehima, and Themeda. [5] [4]
Spikelets within the inflorescence (flower cluster) are generally arranged on spicate racemes in pairs. A fertile, unstalked spikelet is subtended by a sterile, stalked spikelet. In species where awns are present they are found on the fertile, unstalked spikelet as an extension of the lemma. [6]
Classification following Soreng et al. (2017). [1]
The Bambuseae are the most diverse tribe of bamboos in the grass family (Poaceae). They consist of woody species from tropical regions, including some giant bamboos. Their sister group are the small herbaceous bamboos from the tropics in tribe Olyreae, while the temperate woody bamboos (Arundinarieae) are more distantly related. The Bambuseae fall into two clades, corresponding to species from the Neotropics and from the Paleotropics.
Chloridoideae is one of the largest subfamilies of grasses, with roughly 150 genera and 1,600 species, mainly found in arid tropical or subtropical grasslands. Within the PACMAD clade, their sister group is the Danthonioideae. The subfamily includes widespread weeds such as Bermuda grass or goosegrass, but also millet species grown in some tropical regions, namely finger millet and teff.
Panicoideae is the second-largest subfamily of the grasses with over 3,500 species, mainly distributed in warm temperate and tropical regions. It comprises some important agricultural crops, including sugarcane, maize, sorghum, and switchgrass.
Andropogon is a widespread genus of plants in the grass family, native to much of Asia, Africa, and the Americas, as well as Southern Europe and various oceanic islands.
Ischaemum is a genus in the tribe Andropogoneae, belonging to the grass family, widespread in tropical and semitropical regions in many countries. Many species are known commonly as murainagrass.
Oryzeae is a tribe of flowering plants in the true grass family, Poaceae. It contains 11 genera, including both cultivated rice (Oryza) and wild rice (Zizania).
Dimeria is a genus of Asian, Australian, Madagascan, and Pacific Island plants in the grass family, mostly distributed in Peninsular India
The Stipeae are a tribe of grasses within the subfamily Pooidae, with up to 600 described species.
Oryzoideae (syn. Ehrhartoideae) is a subfamily of the true grass family Poaceae. It has around 120 species in 19 genera, notably including the major cereal crop rice. Within the grasses, this subfamily is one of three belonging to the species-rich BOP clade, which all use C3 photosynthesis; it is the basal lineage of the clade.
Paniceae is a large tribe of the subfamily Panicoideae in the grasses (Poaceae), the only in the monotypic supertribe Panicodae. It includes roughly 1,500 species in 84 genera, primarily found in tropical and subtropical regions of the world. Paniceae includes species using either of the C4 and C3 photosynthetic pathways, as well as presumably intermediate species. Most of the millets are members of tribe Paniceae.
Micrairoideae is a subfamily of the grass family Poaceae, distributed in tropical and subtropical regions. Within the PACMAD clade, it is sister to subfamily Arundinoideae.
Phyllorachideae is a tribe in the grass family, comprising two genera. It may be better placed as a subtribe of Oryzeae.
Eriachneae is a tribe of grasses in subfamily Micrairoideae, with 50 species in two genera. Species in the tribe use the C4 photosynthetic pathway and are distributed mainly in Australasia, reaching into Asia and Micronesia.
Paspaleae is a tribe of the Panicoideae subfamily in the grasses (Poaceae), native mainly to the tropical and subtropical Americas but with a number of species introduced to other regions. It includes roughly 680 species in 39 genera. Species in this tribe use either of the C3 or C4 photosynthetic pathways.
Zeugiteae is a tribe of the subfamily Panicoideae in the grasses (Poaceae), native to Africa, Asia, Australasia, South and Central America. There are 18 species in four genera. The tribe belongs to a basal lineage within the subfamily. Species in this tribe use the C3 photosynthetic pathway.
Duthieeae is a tribe of grasses, subfamily Pooideae, containing eight genera.
Ischaemum rugosum, also known as saramollagrass, is a flowering plant belonging to the grass family Poaceae in the genus Ischaemum, and is native to tropical and temperate regions of Asia, growing in marshes and other wet habitats. It is a vigorous annual, and is an invasive species in South America and Madagascar. It reaches heights of up to 1 m and is primarily recognized by the ridged surface of its sessile spikelet’s lower glume. Despite its historic importance as fodder in Asia, the grass has become a major weed in mid-latitude rice paddies throughout Asia and South America.
Rottboellia cochinchinensis is a species of grass known by the common names Itchgrass,Raoul grass, corngrass, Kokoma grass, Guinea-fowl grass, jointed grass, Shamwa grass and Kelly grass. It is a tall, tufted annual grass whose stems (culms) grow up to 3 metres in height with leaf-blades of up to 45 centimetres in length. The species flowers at the apex of culms in the form of spike-like racemes composed of paired spikelets. The common name Itchgrass comes from the bristly (hispid) leaf-sheath which can be irritating to the skin.
Claviceps pusilla, also known as bluestem ergot, is a parasitic fungus primarily of the grass tribe Angropogoneae, particularly those in the tribe referred to as "bluestem". C. pusilla occasionally manifests characteristic triangular conidia which appear to be unique among Claviceps species.
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