The Silurian is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at 443.8 million years ago (Mya), to the beginning of the Devonian Period, 419.2 Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out.
Tentaculites is an extinct genus of conical fossils of uncertain affinity, class Tentaculita, although it is not the only member of the class. It is known from Lower Ordovician to Upper Devonian deposits both as calcitic shells with a brachiopod-like microstructure and carbonaceous 'linings'. The "tentaculites" are also referred to as the styliolinids.
Bioerosion describes the breakdown of hard ocean substrates – and less often terrestrial substrates – by living organisms. Marine bioerosion can be caused by mollusks, polychaete worms, phoronids, sponges, crustaceans, echinoids, and fish; it can occur on coastlines, on coral reefs, and on ships; its mechanisms include biotic boring, drilling, rasping, and scraping. On dry land, bioerosion is typically performed by pioneer plants or plant-like organisms such as lichen, and mostly chemical or mechanical in nature.
Tabulata, commonly known as tabulate corals, are an order of extinct forms of coral. They are almost always colonial, forming colonies of individual hexagonal cells known as corallites defined by a skeleton of calcite, similar in appearance to a honeycomb. Adjacent cells are joined by small pores. Their distinguishing feature is their well-developed horizontal internal partitions (tabulae) within each cell, but reduced or absent vertical internal partitions. They are usually smaller than rugose corals, but vary considerably in shape, from flat to conical to spherical.
Stromatoporoidea is an extinct clade of sea sponges common in the fossil record from the Middle Ordovician to the Late Devonian. They can be characterized by their densely layered calcite skeletons lacking spicules. Stromatoporoids were among the most abundant and important reef-builders of their time, living close together in flat biostromes or elevated bioherms on soft tropical carbonate platforms.
Carbonate hardgrounds are surfaces of synsedimentarily cemented carbonate layers that have been exposed on the seafloor. A hardground is essentially, then, a lithified seafloor. Ancient hardgrounds are found in limestone sequences and distinguished from later-lithified sediments by evidence of exposure to normal marine waters. This evidence can consist of encrusting marine organisms, borings of organisms produced through bioerosion, early marine calcite cements, or extensive surfaces mineralized by iron oxides or calcium phosphates. Modern hardgrounds are usually detected by sounding in shallow water or through remote sensing techniques like side-scan sonar.
Trypanites is a narrow, cylindrical, unbranched boring which is one of the most common trace fossils in hard substrates such as rocks, carbonate hardgrounds and shells. It appears first in the Lower Cambrian, was very prominent in the Ordovician Bioerosion Revolution, and is still commonly formed today. Trypanites is almost always found in calcareous substrates, most likely because the excavating organism used an acid or other chemical agent to dissolve the calcium carbonate. Trypanites is common in the Ordovician and Silurian hardgrounds of Baltica.
Cornulitida is an extinct order of encrusting animals from class Tentaculita, which were common around the globe in the Ordovician to Devonian oceans, and survived until the Carboniferous. Organisms that may be the oldest cornulitids have been found in Cambrian sediments of Jordan.
The order Microconchida is a group of small, spirally-coiled, encrusting fossil "worm" tubes from the class Tentaculita found from the Upper Ordovician to the Middle Jurassic (Bathonian) around the world. They have lamellar calcitic shells, usually with pseudopunctae or punctae and a bulb-like origin. Many were long misidentified as the polychaete annelid Spirorbis until studies of shell microstructure and formation showed significant differences. All pre-Cretaceous "Spirorbis" fossils are now known to be microconchids. Their classification at the phylum level is still debated. Most likely they are some form of lophophorate, a group which includes phoronids, bryozoans and brachiopods. Microconchids may be closely related to the other encrusting tentaculitoid tubeworms, such as Anticalyptraea, trypanoporids and cornulitids.
Tentaculita is an extinct class of uncertain placement ranging from the Early Ordovician to the Middle Jurassic. They were suspension feeders with a near worldwide distribution. For a more thorough discussion, see Tentaculites.
Chaetosalpinx is an ichnogenus of bioclaustrations. Chaetosalpinx includes straight to sinuous cavities that are parallel to the host's axis of growth. The cavity is circular to oval in cross-section and it lacks a wall lining or floor-like tabulae. They are common in tabulate and rugose corals from Late Ordovician to Devonian of Europe and North America. They may have been parasites.
Cornulites is a genus of cornulitid tubeworms. Their shells have vesicular wall structure, and are both externally and internally annulated. They usually occur as encrusters on various shelly fossils. Their fossils are known from the Middle Ordovician to the Carboniferous.
Conchicolites is a fossil genus of cornulitid tubeworms. Their shells lack vesicular wall structure and have a smooth lumen. They are externally covered with transverse ridges. Some species have spines. They usually occur as encrusters on various shelly fossils. Their fossils are known from the Late Ordovician to the Devonian.
Septalites is a genus of cornulitid tubeworms. Their shells lack vesicular wall structure and have a smooth lumen filled with numerous transverse septa. They are externally covered with transverse ridges. Their fossils are known only from the Silurian of Gotland.
Coralloconchus is a genus of cornulitid tubeworms with small, slender, irregularly curved conical tubes with slowly increasing diameter. Tubes have thin walls and a smooth lumen. Tube wall has a lamellar microstructure. Tubes are devoid of septa and vesicles in the adult part and are not spirally coiled.
Punctaconchus is a genus of microconchid tubeworms. It was the last genus of microconchids, and the only genus to exist beyond the Triassic. Their tubes have large pores (punctae) penetrating the tube wall. Tubes lumen is covered by ripplemark−like transverse ridges. Punctaconchus occurs in the Middle Jurassic of England, France and Poland.
Palaeoconchus is a genus of microconchid tubeworms. Their tubes have pseudopunctae penetrating the tube wall. Tubes lumen is smooth. Palaeoconchus occurs in the Late Ordovician of Baltica and Avalonia. In the Devonian it had a global distribution.
Annuliconchus is a genus of microconchid tubeworms. Their tubes have pseudopunctae penetrating the tube wall. Tubes lumen is annulated. Annuliconchus occurs in the Silurian of Baltica.
Tymbochoos is an extinct genus of encrusting tentaculitoid tubeworms. Tymbochoos has a laminar tube structure and pseudopuncta similar to those of the tentaculitoids. It has previously been interpreted as a Palaeozoic polychaete. The world's oldest build-ups with tube-supported frameworks belong to Tymbochoos sinclairi. They occur in the Ordovician limestones of the Ottawa Valley.
Olev Vinn is an Estonian paleobiologist and paleontologist.
