Nematomorpha are a phylum of parasitoid animals superficially similar to nematode worms in morphology, hence the name. Most species range in size from 50 to 100 millimetres, reaching 2 metres (79 in) in extreme cases, and 1 to 3 millimetres in diameter. Horsehair worms can be discovered in damp areas, such as watering troughs, swimming pools, streams, puddles, and cisterns. The adult worms are free-living, but the larvae are parasitic on arthropods, such as beetles, cockroaches, mantises, orthopterans, and crustaceans. About 351 freshwater species are known and a conservative estimate suggests that there may be about 2000 freshwater species worldwide. The name "Gordian" stems from the legendary Gordian knot. This relates to the fact that nematomorphs often coil themselves in tight balls that resemble knots.
Priapulida, sometimes referred to as penis worms, is a phylum of unsegmented marine worms. The name of the phylum relates to the Greek god of fertility, because their general shape and their extensible spiny introvert (eversible) proboscis may resemble the shape of a human penis. They live in the mud and in comparatively shallow waters up to 90 metres (295 ft) deep. Some species show a remarkable tolerance for hydrogen sulfide, anoxia and low salinity. Halicryptus spinulosus appears to prefer brackish shallow waters. They can be quite abundant in some areas. In an Alaskan bay as many as 85 adult individuals of Priapulus caudatus per square meter has been recorded, while the density of its larvae can be as high as 58,000 per square meter.
Hallucigenia is a genus of lobopodian known from Cambrian aged fossils in Burgess Shale-type deposits in Canada and China, and from isolated spines around the world. The generic name reflects the type species' unusual appearance and eccentric history of study; when it was erected as a genus, H. sparsa was reconstructed as an enigmatic animal upside down and back to front. Lobopodians are a grade of Paleozoic panarthropods from which the velvet worms, water bears, and arthropods arose.
The Maotianshan Shales (帽天山页岩) are a series of Early Cambrian sedimentary deposits in the Chiungchussu Formation, famous for their Konservat Lagerstätten, deposits known for the exceptional preservation of fossilized organisms or traces. The Maotianshan Shales form one of some forty Cambrian fossil locations worldwide exhibiting exquisite preservation of rarely preserved, non-mineralized soft tissue, comparable to the fossils of the Burgess Shale of British Columbia, Canada. They take their name from Maotianshan Hill in Chengjiang County, Yunnan Province, China.
Scalidophora is a group of marine pseudocoelomate ecdysozoans that was proposed on morphological grounds to unite three phyla: the Kinorhyncha, the Priapulida and the Loricifera. The three phyla have four characters in common — chitinous cuticle that is moulted, rings of scalids on the introvert, flosculi, and two rings of introvert retracts. The introvert and abdomen are separated by a distinct neck region in all groups, but in adult macroscopic priapulids it becomes rudimentary in Priapulus and is completely absent in Halicryptus. However, the monophyly of the Scalidophora was not supported by two molecular studies, where the position of the Loricifera was uncertain or as sister to the Panarthropoda. Both studies supported a reduced Scalidophora comprising the Kinorhyncha and Priapulida as sister phyla. Their closest relatives are the Panarthropoda, Nematoda and Nematomorpha.
Forfexicaris valida is a species of Lower Cambrian arthropod, the only species in the family Forfexicarididae. It is known from only two specimens from the Maotianshan shale Lagerstätte.
Richard John Aldridge was a British palaeontologist and academic, who was Bennett Professor of Geology at the University of Leicester.
Acanthomeridion is an extinct arthropod found in the Chengjiang fauna deposits of China. In 1997, it was placed in its own, monotypic family, Acanthomeridiidae. It is known from eight specimens, all found in China.
Chengjiang is a city located in Yuxi, Yunnan Province, China, just north of Fuxian Lake.
Archaeopriapulida is a group of priapulid worms known from Cambrian lagerstätte. The group is closely related to, and very similar to, the modern Priapulids. It is unclear whether it is mono- or polyphyletic. Despite a remarkable morphological similarity to their modern cousins, they fall outside of the priapulid crown group, which is not unambiguously represented in the fossil record until the Carboniferous. In addition to well-preserved body fossils, remains of several archaeopriapulid taxa are known to have been preserved primarily as organic microfossils, such as isolated scalids and pharyngeal teeth. They are probably closely related or paraphyletic to the palaeoscolecids; the relationship between these basal worms is somewhat unresolved.
The palaeoscolecids are a group of extinct ecdysozoan worms resembling armoured priapulids. They are known from the Lower Cambrian to the lower Ludfordian ; they are mainly found as disarticulated sclerites, but are also preserved in many of the Cambrian lagerstätten. They take their name from the typifying genus Palaeoscolex. Other genera include Cricocosmia from the Lower Cambrian Chengjiang biota. Their taxonomic affinities within Ecdysozoa have been the subject of debate.
Maotianshania cylindrica is an extinct worm-like creature of average size. It occurs in the Lower Cambrian (Atdabanian) Chengjiang biota of Northeastern Yunnan, China. It is usually preserved as pink impression. The gut is a dark central zone. The anterior pharnyx and "collar", carry tiny sepia-colored teeth. The very back-end of the body has two small hooks.
Cotyledion tylodes is an extinct, stalked filter-feeder known from the Chengjiang lagerstatten. The living animal reached a couple of centimetres in height, and bore a loose scleritome of ovoid sclerites. Its interpretation has been controversial, but it is currently thought to be a member of the Entoprocta stem group.
Acosmia is an extinct genus of marine worm from the Cambrian aged Chengjiang biota of Yunnan, China. It is represented by a single rare species, Acosmia maotiania, that reached 45 mm in length and 9 mm in width. It was likely a burrowing animal that fed by deposit feeding. While originally suggested to be a priapulid, a 2020 study proposed to be a stem-group ecdysozoan, due to lacking the radial pharygneal armature that characterises modern ecdysozoans, including priapulids.
Paraselkirkia is a genus of archaeopriapulid known from the Chengjiang biota, resembling Selkirkia.
Strabops is a genus of strabopid, an extinct group of arthropods. Strabops is known from a single specimen from the Late Cambrian of the Potosi Dolomite, Missouri, collected by a former professor, Arthur Thacher. It is classified in the family Strabopidae of the monotypic order Strabopida, a group closely related to the aglaspidids with uncertain affinities. The generic name is composed by the Ancient Greek words στραβός, meaning "squinting", and ὄψῐς, meaning "face".
Fuxianospira gyrata is a Cambrian macroalgae found in the Chengjiang lagerstätte. Preserved in clustered, helicoid groups, the filaments are threadlike, plain and without branches. Brown and smooth in appearance, these structural characteristics display a resemblance to modern brown algae. A limited amount of algae species have been discovered in the Chengjiang biota, suggesting that diversity within the general algae population may have been sparse. It is thought that the macroalgae could be the most basic component of the Chengjiang biota food chain.
Paragordius is a genus of worms belonging to the family Chordodidae. It was independently described by both Lorenzo Camerano in 1897 and Thomas Harrison Montgomery Jr. in 1898, though both authors gave the genus the same name.
Vetulocystis is an extinct genus type genus of the family Vetulocystidae. It was found in the Maotianshan Shales, which date to roughly 518 mya.
Hallucigeniidae is a family of extinct worms belonging to the group Lobopodia that originated during the Cambrian explosion. It is based on the species Hallucigenia sparsa, the fossil of which was discovered by Charles Doolittle Walcott in 1911 from the Burgess Shale of British Columbia. The name Hallucigenia was created by Simon Conway Morris in 1977, from which the family was erected after discoveries of other hallucigeniid worms from other parts of the world. Classification of these lobopods and their relatives are still controversial, and the family consists of at least four genera.
