Arcillas de Morella Formation | |
---|---|
Stratigraphic range: | |
Type | Geological formation |
Underlies | Xert Formation |
Overlies | Artoles Formation |
Lithology | |
Primary | Sandstone, clay |
Other | Marl, limestone, siltstone, conglomerate |
Location | |
Region | Europe |
Country | Spain |
Type section | |
Named for | Morella, Castellón |
Named by | Canérot et al. |
Year defined | 1982 |
The Arcillas de Morella Formation is a geological formation in Spain whose strata date back to the Barremian stage of the Early Cretaceous. [1] Dinosaur remains are among the fossils that have been recovered from the formation. [2]
The formation was formally named and defined by Canérot and colleagues in 1982. [3] The formation predominantly consists of red clays and continental sandstones, with some marine intercalations. [4]
Dinosaurs of the Arcillas de Morella Formation | |||
---|---|---|---|
Taxa | Species | Notes | Images |
Hypsilophodon [2] | |||
Iguanodon | I. bernissartensis [5] | An iguanodonian dinosaur | |
Mantellisaurus | M. cf. atherfieldensis [5] | An iguanodontian dinosaur | |
Morelladon | Morelladon beltrani [6] | An iguanodontian dinosaur | |
Vallibonavenatrix | V. cani [4] | A spinosaurid theropod | |
Protathlitis | P. cinctorrensis [7] | A spinosaurid theropod | |
Baryonychinae [2] | Indeterminate remains. [2] | ||
Garumbatitan | G. morellensis | A somphospondylan sauropod [8] | |
Titanosauriformes | Indeterminate | At least two distinct taxa, including a somphospondylan [9] | |
Turtles of the Arcillas de Morella Formation | |||
---|---|---|---|
Taxa | Species | Notes | Images |
Brodiechelys | B. royoi [10] | A xinjiangchelyid turtle | |
Eodortoka | E. morellana [11] | A dortokid turtle |
Plesiosaurs of the Arcillas de Morella Formation | ||||
---|---|---|---|---|
Taxa | Presence | Notes | Images | |
Leptocleididae [12] |
Baryonyx is a genus of theropod dinosaur which lived in the Barremian stage of the Early Cretaceous period, about 130–125 million years ago. The first skeleton was discovered in 1983 in the Smokejack Clay Pit, of Surrey, England, in sediments of the Weald Clay Formation, and became the holotype specimen of Baryonyx walkeri, named by palaeontologists Alan J. Charig and Angela C. Milner in 1986. The generic name, Baryonyx, means "heavy claw" and alludes to the animal's very large claw on the first finger; the specific name, walkeri, refers to its discoverer, amateur fossil collector William J. Walker. The holotype specimen is one of the most complete theropod skeletons from the UK, and its discovery attracted media attention. Specimens later discovered in other parts of the United Kingdom and Iberia have also been assigned to the genus, though many have since been moved to new genera.
The Spinosauridae are a clade or family of tetanuran theropod dinosaurs comprising ten to seventeen known genera. Spinosaurid fossils have been recovered worldwide, including Africa, Europe, South America and Asia. Their remains have generally been attributed to the Early to Mid Cretaceous.
Siamosaurus is a genus of spinosaurid dinosaur that lived in what is now known as China and Thailand during the Early Cretaceous period and is the first reported spinosaurid from Asia. It is confidently known only from tooth fossils; the first were found in the Sao Khua Formation, with more teeth later recovered from the younger Khok Kruat Formation. The only species Siamosaurus suteethorni, whose name honours Thai palaeontologist Varavudh Suteethorn, was formally described in 1986. In 2009, four teeth from China previously attributed to a pliosaur—under the species "Sinopliosaurus" fusuiensis—were identified as those of a spinosaurid, possibly Siamosaurus. It is yet to be determined if two partial spinosaurid skeletons from Thailand and an isolated tooth from Japan also belong to Siamosaurus.
Mantellisaurus is a genus of iguanodontian dinosaur that lived in the Barremian and early Aptian ages of the Early Cretaceous Period of Europe. Its remains are known from Belgium (Bernissart), England, Spain and Germany. The type and only species is M. atherfieldensis. Formerly known as Iguanodon atherfieldensis, the new genus Mantellisaurus was erected for the species by Gregory Paul in 2007. According to Paul, Mantellisaurus was more lightly built than Iguanodon and more closely related to Ouranosaurus, making Iguanodon in its traditional sense paraphyletic. It is known from many complete and almost complete skeletons. The genus name honours Gideon Mantell, the discoverer of Iguanodon.
The Lourinhã Formation is a fossil rich geological formation in western Portugal, named for the municipality of Lourinhã. The formation is mostly Late Jurassic in age (Kimmeridgian/Tithonian), with the top of the formation extending into the earliest Cretaceous (Berriasian). It is notable for containing a fauna especially similar to that of the Morrison Formation in the United States and a lesser extent to the Tendaguru Formation in Tanzania. There are also similarities to the nearby Villar del Arzobispo Formation and Alcobaça Formation. The stratigraphy of the formation and the basin in general is complex and controversial, with the constituent member beds belonging to the formation varying between different authors.
The Blesa Formation is a geological formation in Teruel and La Rioja, Spain whose strata date back to the Barremian of the Early Cretaceous. Dinosaur remains as well as eggs are among the fossils that have been recovered from the formation. Along with the fragmentary anhanguerian pterosaur Iberodactylus. While the lower and upper parts of the formation were deposited in a continental setting, the middle portion of the formation is largely nearshore marine, with remains of plesiosaurs.
The La Huérguina Formation is a geological formation in Spain whose strata date back to the Barremian stage of the Early Cretaceous. Las Hoyas is a Konservat-Lagerstätte within the formation, located near the city of Cuenca, Spain. The site is mostly known for its exquisitely preserved dinosaurs, especially enantiornithines. The lithology of the formation mostly consists of lacustrine limestone deposited in a freshwater wetland environment.
The Villalba de la Sierra Formation is a Campanian to Maastrichtian geologic formation in Spain. Fossil dinosaur eggs have been reported from the formation, that comprises gypsiferous, grey, argillaceous mudstones and sandstones, deposited in a floodplain environment characterised by high seasonality and variability in water availability.
Concavenator is an extinct allosaurinae|allosaurid]] theropod dinosaur that lived approximately 130 million years ago during the Early Cretaceous period. The type species is C. corcovatus. Concavenator corcovatus means "Cuenca hunter with a hump". The fossil was discovered in the Las Hoyas fossil site of Spain by paleontologists José Luis Sanz, Francisco Ortega, and Fernando Escaso from the Autonomous University of Madrid and the National University of Distance Education.
Ichthyovenator is a genus of spinosaurid dinosaur that lived in what is now Laos, sometime between 125 and 113 million years ago, during the Aptian stage of the Early Cretaceous period. It is known from fossils collected from the Grès supérieurs Formation of the Savannakhet Basin, the first of which were found in 2010, consisting of a partial skeleton without the skull or limbs. This specimen became the holotype of the new genus and species Ichthyovenator laosensis, and was described by palaeontologist Ronan Allain and colleagues in 2012. The generic name, meaning "fish hunter", refers to its assumed piscivorous lifestyle, while the specific name alludes to the country of Laos. In 2014, it was announced that more remains from the dig site had been recovered; these fossils included teeth, more vertebrae (backbones) and a pubic bone from the same individual.
Ostafrikasaurus is a genus of theropod dinosaur from the Late Jurassic period of what is now Lindi Region, Tanzania. It is known only from fossil teeth discovered sometime between 1909 and 1912, during an expedition to the Tendaguru Formation by the Natural History Museum of Berlin. Eight teeth were originally attributed to the dubious dinosaur genus Labrosaurus, and later to Ceratosaurus, both known from the North American Morrison Formation. Subsequent studies attributed two of these teeth to a spinosaurid dinosaur, and in 2012, Ostafrikasaurus crassiserratus was named by French palaeontologist Eric Buffetaut, with one tooth as the holotype, and the other referred to the same species. The generic name comes from the German word for German East Africa, the former name of the colony in which the fossils were found, while the specific name comes from the Latin words for "thick" and "serrated", in reference to the form of the animal's teeth.
Camarillasaurus is a genus of spinosaurid dinosaur from the Early Cretaceous period (Barremian) of Camarillas, Teruel Province, in what is now northeastern Spain. Described in 2014, it was originally identified as a ceratosaurian theropod, before being referred to the Spinosauridae in 2021.
Morelladon is an extinct genus of herbivorous styracosternan ornithopod dinosaur. It lived during the Early Cretaceous of Spain, around 130 million years ago.
Vallibonavenatrix is a genus of spinosaurid dinosaur from the Early Cretaceous (Barremian) Arcillas de Morella Formation of Castellón, Spain. The type and only species is Vallibonavenatrix cani, known from a partial skeleton.
Dortokidae is an extinct family of freshwater pan-pleurodiran turtles, known from the Cretaceous and Paleocene of Europe. Only four species have been named, but indeterminate fossils show that they were abundant across western and eastern-central Europe during the Cretaceous. The family is only known from postcranial remains.
Portellsaurus is a genus of hadrosauroid dinosaur from the Lower Cretaceous Margas de Mirambell Formation of Spain. The genus contains a single species, Portellsaurus sosbaynati, known from a partial right dentary.
Baryonychinae is an extinct clade or subfamily of spinosaurids from the Early Cretaceous (Valanginian-Albian) of Britain, Portugal, and Niger. In 2023, it consisted of seven genera: Ceratosuchops, Cristatusaurus, Protathlitis, Riparovenator, Suchomimus, Suchosaurus, and Baryonyx, the nominal genus. The clade was named by Charig & Milner in 1986 and defined by Sereno et al. in 1998 and Holtz et al. in 2004 as all taxa more closely related to Baryonyx walkeri than to Spinosaurus aegyptiacus.
Protathlitis is a genus of spinosaurid dinosaur from the Early Cretaceous (Barremian) Arcillas de Morella Formation of Castellón, Spain. The type and only species is Protathlitis cinctorrensis, known from a partial skeleton. It was a basal member of the Baryonychinae. Its discovery, as well as that of the spinosaurine Vallibonavenatrix shows that the Iberian Peninsula held a diverse assemblage of spinosaurids during the Early Cretaceous.
Garumbatitan is an extinct genus of somphospondylan sauropod dinosaur from the Cretaceous Arcillas de Morella Formation of Spain. The genus contains a single species, G. morellensis, known from multiple partial skeletons.