Barylambdidae | |
---|---|
Barylambda faberi | |
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | † Cimolesta |
Suborder: | † Pantodonta |
Superfamily: | † Pantolambdoidea |
Family: | † Barylambdidae Patterson, 1939 |
Genera | |
Barylambdidae is an extinct family of pantodont mammals from North America. [1]
Odd-toed ungulates, mammals which constitute the taxonomic order Perissodactyla, are animals—ungulates—who have reduced the weight-bearing toes to three or one of the five original toes. The non-weight-bearing toes are either present, absent, vestigial, or positioned posteriorly. By contrast, the even-toed ungulates bear most of their weight equally on four or two of the five toes: their third and fourth toes. Another difference between the two is that odd-toed ungulates digest plant cellulose in their intestines, rather than in one or more stomach chambers as even-toed ungulates, with the exception of Suina, do.
Ungulates are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. Living ungulates are divided into two orders: the odd-toed ungulates (Perissodactyla) including horses, rhinoceroses, and tapirs; and even-toed ungulates (Artiodactyla) such as cattle, pigs, giraffes, camels, sheep, deer, and hippopotamuses. Cetaceans such as whales, dolphins, and porpoises are also classified as even-toed ungulates, although they do not have hooves. Most terrestrial ungulates use the hoofed tips of their toes to support their body weight while standing or moving. Two other orders of ungulates, Notoungulata and Litopterna, both native to South America, became extinct at the end of the Pleistocene, around 12,000 years ago.
The even-toed ungulates are ungulates—hoofed animals—which bear weight equally on two of their five toes: the third and fourth. The other three toes are either present, absent, vestigial, or pointing posteriorly. By contrast, odd-toed ungulates bear weight on an odd number of the five toes. Another difference between the two is that many other even-toed ungulates digest plant cellulose in one or more stomach chambers rather than in their intestine as the odd-toed ungulates do.
Camelids are members of the biological family Camelidae, the only currently living family in the suborder Tylopoda. The seven extant members of this group are: dromedary camels, Bactrian camels, wild Bactrian camels, llamas, alpacas, vicuñas, and guanacos. Camelids are even-toed ungulates classified in the order Cetartiodactyla, along with species like whales, pigs, deer, cattle, and antelopes.
Litopterna is an extinct order of South American native ungulates. Like other endemic South American mammals, their relationship to other mammal groups had long been unclear, but recent genetic and proteomic evidence indicates that their closest living relatives are Perissodactyls including horses, rhinoceros, and tapirs, and that litopterns are closely related to notoungulates, another widespread group of South American ungulates.
Condylarthra is an informal group – previously considered an order – of extinct placental mammals, known primarily from the Paleocene and Eocene epochs. They are considered early, primitive ungulates. It is now largely considered to be a wastebasket taxon, having served as a dumping ground for classifying ungulates which had not been clearly established as part of either Perissodactyla or Artiodactyla, being composed thus of several unrelated lineages.
Notoungulata is an extinct order of mammalian ungulates that inhabited South America from the early Paleocene to the Holocene, living from approximately 61 million to 11,000 years ago. Notoungulates were morphologically diverse, with forms resembling animals as disparate as rabbits and rhinoceroses. Notoungulata are the largest group of South American native ungulates, with over 150 genera in 14 families having been described, divided into two major subgroupings, Typotheria and Toxodontia. Notoungulates first diversified during the Eocene. Their diversity declined during the Late Neogene, with only the large toxodontids persisting until the end of the Pleistocene. Collagen analysis suggests that notoungulates are closely related to litopterns, another group of South American ungulates, and their closest living relatives being perissodactyls, including rhinoceroses, tapirs and equines. However their relationships to other South American ungulates are uncertain. Several groups of notoungulates separately evolved ever-growing cheek teeth.
The Barbary sheep, also known as aoudad is a species of caprine native to rocky mountains in North Africa. While this is the only species in genus Ammotragus, six subspecies have been described. Although it is rare in its native North Africa, it has been introduced to North America, southern Europe, and elsewhere. It is also known in the Berber language as waddan or arwi, and in former French territories as the moufflon.
South American native ungulates, commonly abbreviated as SANUs, are extinct ungulate-like mammals of controversial affinities that were indigenous to South America prior to the Great American Biotic Interchange. They comprise five major groups conventionally ranked as orders—Astrapotheria, Litopterna, Notoungulata, Pyrotheria, and Xenungulata—as well as the primitive "condylarth" groups Didolodontidae and Kollpaniinae. It has been proposed that some or all of the members of this group form a clade, named Meridiungulata, though the relationships of South American ungulates remain largely unresolved. The two largest groups of South American ungulates, the notoungulates and the litopterns, were the only groups to persist beyond the mid Miocene. Only a few of the largest species of notoungulates and litopterns survived until the end-Pleistocene extinctions.
Pantodonta is an extinct suborder of eutherian mammals. These herbivorous mammals were one of the first groups of large mammals to evolve after the end of the Cretaceous. The last pantodonts died out at the end of the Eocene.
Mesonychidae is an extinct family of small to large-sized omnivorous-carnivorous mammals. They were endemic to North America and Eurasia during the Early Paleocene to the Early Oligocene, and were the earliest group of large carnivorous mammals in Asia. They are not closely related to any living mammals. Mesonychid taxonomy has long been disputed and they have captured popular imagination as "wolves on hooves," animals that combine features of both ungulates and carnivores. Skulls and teeth have similar features to early whales, and the family was long thought to be the ancestors of cetaceans. Recent fossil discoveries have overturned this idea; the consensus is that whales are highly derived artiodactyls. Some researchers now consider the family a sister group either to whales or to artiodactyls, close relatives rather than direct ancestors. Other studies define Mesonychia as basal to all ungulates, occupying a position between Perissodactyla and Ferae. In this case, the resemblances to early whales would be due to convergent evolution among ungulate-like herbivores that developed adaptations related to hunting or eating meat.
Toxodontidae is an extinct family of notoungulate mammals, known from the Oligocene to the Holocene of South America, with one genus, Mixotoxodon, also known from the Pleistocene of Central America and southern North America. Member of the family were medium to large-sized, and had medium to high-crowned dentition, which in derived members of the group evolved into ever-growing cheek teeth. Isotopic analyses have led to the conclusion that Pleistocene members of the family were mixed feeders.
Cormocyon is an extinct genus of borophagine canid native to North America. It lived from the Oligocene to the Early Miocene, 30.8—20.6 Mya, existing for about 10.2 million years. It is regarded as a primitive, transitional member of the Borophagini tribe.
Ursavini is an extinct tribe of mammals of the family Ursidae (bears) endemic to North America, Europe, Africa, and Asia during Miocene through Pliocene, living from about 23—2.5 Mya, existing for roughly 20.5 million years.
Phenacodontidae is an extinct family of large herbivorous mammals traditionally placed in the “wastebasket taxon” Condylarthra, which may instead represent early-stage perissodactyls. They lived in the Paleocene and Eocene epochs and their fossil remains have been found in North America and Europe.
Didolodontidae is a possibly paraphyletic family of "condylarth" mammals known from the Paleogene of South America, with most specimens known from Argentina. They were generally small-medium in body size, and had a bunodont dentition. A close relationship with litopterns has been suggested by some studies. They range in age from the early Paleocene (Selandian/Peligran) to late Eocene (Priabonian/Mustersan). The attribution of Salladolodus deuterotheroides from the Late Oligocene of Bolivia to the family is doubtful.
Lambertocyon is a genus of ungulates from western North America. Three species are known, making their last appearance in the Late Paleocene Clarkforkian stage.
Zhelestidae is a lineage of extinct eutherian mammals. Occurring in the Late Cretaceous from the Turonian to the Maastrichtian, they were an extremely successful group, with representatives present in Europe, Asia, India, Africa and North America, ostensibly rendering them a cosmopolitan clade. They were specialised towards an herbivorous lifestyle and were in fact initially considered stem-ungulates, but the presence of epipubics and "archaic" dental characters render them as non-placental eutherians.
Hemipsalodon is an extinct genus of hyainailourid hyaenodonts from paraphyletic subfamily Hyainailourinae that lived in North America during the middle to late Eocene.
Molinodus is an extinct genus of mammal of uncertain affinities, whose fossils were found at Tiupampa, in Bolivia, in terrains dated from the Paleocene.