Barylambda | |
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B. faberi skeleton, Field Museum | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | † Pantodonta |
Family: | † Barylambdidae |
Genus: | † Barylambda Patterson 1937 |
Species [1] | |
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Barylambda (Greek: "heavy" (baros), "lambda" (lambda) in a reference to larger size than that of Pantolambda [2] ) is an extinct genus of pantodont mammal from the middle to late Paleocene, well known from several finds in the Wasatchian (NALMA classification) DeBeque Formation of Colorado and the Clarkforkian Wasatch Formation to Tiffanian Fort Union Formation in Wyoming. Three species of Barylambda are currently recognized. [3] The creature likely lived a life similar to that of a modern tapir, browsing on foliage and soft vegetation. [4] Barylambda seems to have been quite successful for an early pantodont, though eventually it seems to have been replaced in its ecosystem by other pantodonts, such as Coryphodon .
In life, Barylambda probably resembled a large ground sloth, with a small head and long, well-developed tail and bear-like legs. It was a large animal weighing around 650 kg (1,430 lb), [5] about the size of a pony. Barylambda was large even for a pantodont, sheer size probably protecting it from contemporary carnivores.
Like other pantodonts, Barylambda was a heavyset, five-toed plantigrade animal. The vertebrae of the tail were unusually massive; the living animal may have been able to rear up and support itself on the hind legs and tail in order to reach higher for food. The generalized appearance of the teeth, the presence of well-developed canines only in males, the grinding wear and lack of shearing blades on the molars, and the animal's heavy build strongly suggest that it was herbivorous.
Ungulates are members of the diverse clade Euungulata, which primarily consists of large mammals with hooves. Once part of the clade "Ungulata" along with the clade Paenungulata, "Ungulata" has since been determined to be a polyphyletic and thereby invalid clade based on molecular data. As a result, true ungulates had since been reclassified to the newer clade Euungulata in 2001 within the clade Laurasiatheria while Paenungulata has been reclassified to a distant clade Afrotheria. Living ungulates are divided into two orders: Perissodactyla including equines, rhinoceroses, and tapirs; and Artiodactyla including cattle, antelope, pigs, giraffes, camels, sheep, deer, and hippopotamuses, among others. Cetaceans such as whales, dolphins, and porpoises are also classified as artiodactyls, although they do not have hooves. Most terrestrial ungulates use the hoofed tips of their toes to support their body weight while standing or moving. Two other orders of ungulates, Notoungulata and Litopterna, both native to South America, became extinct at the end of the Pleistocene, around 12,000 years ago.
Creodonta is a former order of extinct carnivorous placental mammals that lived from the early Paleocene to the late Miocene epochs in North America, Europe, Asia and Africa. Originally thought to be a single group of animals ancestral to the modern Carnivora, this order is now usually considered a polyphyletic assemblage of two different groups, the oxyaenids and the hyaenodontids, not a natural group. Oxyaenids are first known from the Palaeocene of North America, while hyaenodonts hail from the Palaeocene of Africa.
Chapalmalania is an extinct genus of procyonid from the Late Miocene or Early Pliocene to Early Pleistocene of Argentina, Venezuela, and Colombia.
Chriacus is an extinct genus of placental mammals that lived in what is now North America during the Paleocene epoch and died out after the early Eocene. In life, members of the genus would have looked something like a kinkajou or binturong, though they were not closely related to any living mammal. Well preserved fossils allow clear information on what they looked like. They were about 1 metre (3.3 ft) long including a long, robust tail, which may or may not have been prehensile. Other features include a light build, weighing approximately 7 kg (15 lb), and many adaptations typical of animals that live in trees. These include walking on the soles of their five-toed feet, and having long, curved, compressed claws. The powerfully built limbs had flexible joints, especially the ankles, an adaptation that allows an animal to turn its hind feet behind it, like modern tree squirrels, in order to climb downward. They were probably omnivores, eating fruit, eggs, insects and small mammals.
Coryphodon is an extinct genus of pantodonts of the family Coryphodontidae.
Didolodus is an extinct genus of mammals from Middle Eocene Argentina. It is an ungulate mammal of uncertain affinities, possibly related to Litopterna, though this is uncertain due to the lack of reliable post-cranial remains, and for now remains Meridiungulata incertae sedis. Its remains were found in the Sarmiento Formation of Patagonia.
Arctocyon is an extinct genus of ungulate mammals. Arctocyon was a "ground dwelling omnivore", that lived from 61.3-56.8 Ma. Synonyms of Arctocyon include Claenodon, and Neoclaenodon. Arctocyon was likely plantigrade, meaning that it walked with its feet flat on the ground, rather than on its toes.
Oxyaena is an extinct genus of placental mammals from extinct subfamily Oxyaeninae within extinct family Oxyaenidae, that lived in Europe, Asia and North America during the early Eocene.
Pantolambda is an extinct genus of Paleocene pantodont mammal. Pantolambda lived during the middle Paleocene, and has been found both in Asia and North America.
Pantodonta is an extinct suborder of eutherian mammals. These herbivorous mammals were one of the first groups of large mammals to evolve after the end of the Cretaceous. The last pantodonts died out at the end of the Eocene.
Arctocyonidae is as an extinct family of unspecialized, primitive mammals with more than 20 genera. Animals assigned to this family were most abundant during the Paleocene, but extant from the late Cretaceous to the early Eocene . Like most early mammals, their actual relationships are very difficult to resolve. No Paleocene fossil has been unambiguously assigned to any living order of placental mammals, and many genera resemble each other: generalized robust, not very agile animals with long tails and all-purpose chewing teeth, living in warm closed-canopy forests with many niches left vacant by the K-T extinction.
Tillodontia is an extinct suborder of eutherian mammals known from the Early Paleocene to Late Eocene of China, the Late Paleocene to Middle Eocene of North America where they display their maximum species diversity, the Middle Eocene of Pakistan, and the Early Eocene of Europe. Leaving no descendants, they are most closely related to the pantodonts, another extinct group. The tillodonts were medium- to large-sized animals that probably fed on roots and tubers in temperate to subtropical habitats.
The Clarkforkian North American Stage, on the geologic timescale, is the North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), typically set from 56,800,000 to 55,400,000 years BP lasting 1.4 million years.
Phenacodontidae is an extinct family of large herbivorous mammals traditionally placed in the “wastebasket taxon” Condylarthra, which may instead represent early-stage perissodactyls. They lived from the late early Paleocene to early middle Eocene and their fossil remains have been found in North America and Europe. The only unequivocal Asian phenacodontid is Lophocion asiaticus.
Riostegotherium was an extinct genus of armadillo from the Early to Late Eocene of South America. Fossils of the genus have been found in the Andesitas Huancache Formation of Argentina and in the Itaboraí Formation, Rio de Janeiro, Brazil.
Alcidedorbignya is an extinct pantodont mammal known from the Early Paleocene Santa Lucia Formation at Tiupampa near Mizque, Cochabamba, Bolivia.
Coryphodontidae is an extinct family of pantodont mammals known from the Late Paleocene to the Middle Eocene of Eurasia and North America.
Protictis is an extinct paraphyletic genus of placental mammals from extinct subfamily Didymictinae within extinct family Viverravidae, that lived in North America from early Paleocene to middle Eocene.
Bemalambda is a genus of extinct mammal, belonging to the pantodonts. It lived in the lower-middle Paleocene and the fossil remains have been found in China.
Periptychus is an extinct genus of mammal belonging to the family Periptychidae. It lived from the Early to Late Paleocene and its fossil remains have been found in North America.