Bellerophontida Temporal range: Anisian records) (Possible | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Mollusca |
Class: | Gastropoda (?) |
Order: | † Bellerophontida Ulrich & Scofield, 1897 |
The Bellerophontida is a taxonomic order of extinct marine mollusks that are found in the fossil record from the Lower Cambrian to the Lower Triassic. They are considered by some experts to be primitive sea snails with primarily symmetrically coiled shells, marine gastropod mollusks.
R.C Moore [1] and J. Brooks Knight et al 1960 [2] recognized bellerophonts as true gastropods, placing them as the suborder Bellerophontina in the subclass Prosobranchia, order Archaeogastropoda. Some more recent workers in the field have expressed uncertainty as to the taxonomic placement of this group.
Shells of bellerophontids are most commonly isotrophic (coiled symmetrically), rarely slightly asymmetric. Most are closely coiled with some general more openly coiled, cyrtiform; predominantly with a median slit in the upper lip of the aperture, or with a series of openings known as tremata, usually generating a selenizone which was probably exhalant in function. The shell wall is of variable thickness and no evidence of nacre or of an operculum has been found. Nothing is known of soft part anatomy of the group, although a simple pair of retractor muscles is inferred from muscle scars. Isotrophic symmetry also suggests paired and equal nephridia and ctenidia (kidneys and gills). [2]
According to Knight, et al, [2] the Bellerophontina, as then regarded, includes two superfamilies, the Helcionellacea, and the Bellerphonitacea. The Helcionellacea are the more primitive, with more cap-like shells, that are limited to the Cambrian Period. Two small families, the Helcionellidae and Coreospiridae are included. The Bellerophontacea, which range from the Upper Cambrian to the Lower Triassic, are more developed and more diverse; that taxon contains three families, the Cyrtolitidae, Sinuitidae, and Bellerophontidae, along with subfamilies and tribes.
Ponder & Lindberg (1997) included the Bellerophontida, as the Bellerophontinaka, in with the Mimophserina, in the Class Gastropoda, incertæ sedis, excluding them from the Eogastropoda, or Prosobranchia. Bouchet & Rocroi (2005) expressed greater uncertainty, placing the bellerophonts, as Bellerophontoidea in Paleozoic molluscs with isostrophically coiled shells of uncertain position (Gastropoda or Monoplacophora).
More recently, J.P. Wager, 2011 [3] included the Bellerophontida as an order in the Gastropoda.
Gastropods, commonly known as slugs and snails, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda.
Monoplacophora, meaning "bearing one plate", is a polyphyletic superclass of molluscs with a cap-like shell, inhabiting deep sea environments. Extant representatives were not recognized as such until 1952; previously they were known only from the fossil record, and were thought to have become extinct 375 million years ago.
Nautiloids are a group of marine cephalopods (Mollusca) which originated in the Late Cambrian and are represented today by the living Nautilus and Allonautilus. Fossil nautiloids are diverse and species rich, with over 2,500 recorded species. They flourished during the early Paleozoic era, when they constituted the main predatory animals. Early in their evolution, nautiloids developed an extraordinary diversity of shell shapes, including coiled morphologies and giant straight-shelled forms (orthocones). No orthoconic and only a handful of coiled species, the nautiluses, survive to the present day.
Bellerophon is a genus of extinct paleozoic marine molluscs of uncertain position in the family Bellerophontidae.
Strepsodiscus is an extinct genus of very primitive fossil snail-like molluscs from the early part of the Late Cambrian of North America. The coiled, slightly asymmetrical shells are about 3 cm in height. It is not known whether these are shells of gastropods or monoplacophorans, which are more primitive mollusks.
The Bellerophontidae are an extinct family of specialized globose bellerophontids, Paleozoic and early Triassic mollusks of the class Gastropoda.
The Euomphalina comprise a major suborder of mainly Paleozoic archaeogastropods, shells of which are hyperstophic to depressed orthstrophic, commonly with an angulation at the outer upper whorl surface thought to be coincident with the exhalent channel; shell wall thick, outer layer calcitic, inner layers aragonitic but not nacreous; operculum calcareous and heavy. Their range is from the Upper Cambrian to the Triassic, and possibly as high as the Upper Cretaceous.
Helcionellid or Helcionelliformes is an order of small fossil shells that are universally interpreted as molluscs, though no sources spell out why this taxonomic interpretation is preferred. These animals are first found about 540 to 530 million years ago in the late Nemakit-Daldynian age, which is the earliest part of the Cambrian period. A single species persisted to the Early Ordovician. These fossils are component of the small shelly fossils (SSF) assemblages.
Helcionelloida is an extinct group of ancient molluscs. These are the oldest known conchiferan molluscs, that is, they had a mineralised shell. Some members of this class were mistaken for Monoplacophorans. The class was erected by Peel in 1991.
Bellerophontoidea, common name "bellerophonts", is a superfamily of extinct planospirally-coiled globose molluscs. This superfamily is generally included within the Gastropoda, but may instead be a group of monoplacophorans. The taxon first appeared late in the Cambrian and continued until late in the Triassic.
Pleurotomarioidea is a superfamily of small to large marine gastropods included in the order Pleurotomariida of the subclass Vetigastropoda.
Bucanellidae is an extinct family of Paleozoic molluscs of uncertain position, belonging either to Gastropoda (snails) or Monoplacophora. The family lived from the upper Cambrian to middle Permian and the shells are characterized by a relatively small median sinus in the upper margin of the aperture, and collabral (transverse) or spiral (longitudinal) threads covering the shell. The shells are planispirally coiled rather than trochospirally with a spire as is the case with most shelled gastropods.
Bucaniidae is an extinct family of Paleozoic molluscs of uncertain position possibly being either gastropods or monoplacophorans in the superfamily Bellerophontoidea. The family lived from the Lower Ordovician to the Devonian and have shells in which the apertural margins tend to flare. Most genera have a slit and selenizone, others some modification of this feature.
Macluritidae is an extinct family of relatively large, Lower Ordovician to Devonian, macluritacean gastropods(?), hypserstrophically coiled, that is dextral while appearing sinistral, of which the genus Maclurites is arch-typical. The base of their shells is flat or gently protruding while the upper side is generally concave.
The †Raphistomatidae is a taxonomic family of fossil sea snails, Paleozoic marine gastropod molluscs. This family was established by Koken in 1896. This family is found in the fossil record from the Upper Cambrian to the Middle Permian.
†Onychochilidae is an extinct family of small, fossil, upper Cambrian to lower Devonian molluscs. They have hyperstrophically coiled shells which generally have smooth whorls and elongate apertures. They are commonly considered to be gastropods, i.e. sea snails.
Euomphaloidea, originally Euomphalacea, is an extinct superfamily of marine molluscs that lived from the Early Ordovician to the Late Cretaceous, included in the Gastropoda but speculated as instead perhaps Monoplacophora.
The Kirengellids are a group of problematic Cambrian fossil shells of marine organisms. The shells bear a number of paired muscle scars on the inner surface of the valve.
Rhynchonelliformea is a major subphylum and clade of brachiopods. It is roughly equivalent to the former class Articulata, which was used previously in brachiopod taxonomy up until the 1990s. These so-called articulated brachiopods have many anatomical differences relative to "inarticulate" brachiopods of the subphyla Linguliformea and Craniformea. Articulates have hard calcium carbonate shells with tongue-and-groove hinge articulations and separate sets of simple opening and closing muscles.
The Trochina is a taxon that is used by paleontologists. It is a suborder of primitive sea snails, marine gastropod mollusks.