Brooksella Temporal range: | |
---|---|
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Porifera |
Class: | Hexactinellida |
Order: | † Reticulosa |
Family: | † Protospongiidae |
Genus: | † Brooksella Walcott, 1896 |
Type species | |
†Brooksella alternata Walcott, 1896 |
Brooksella is an enigmatic star-shaped Cambrian fossil found in the Conasauga Formation of Alabama and Georgia. [1] These fossils are often referred to as "star-cobbles" for their distinct lobate appearance, generally with 6 or more lobes.
Brooksella was first described in 1896 by Charles Doolittle Walcott, who believed them to be medusoid body fossils of cnidarians. [2] [3] Later researchers have offered other explanations, from diagenetic gas bubbles to burrow traces. The most accepted identity is that they are hexactinellid sponges, based on observed spicules, ostia, and internal structure. [1] In 2023, a group of researchers suggested that Brooksella is a pseudofossil, finding no support for previous interpretations of it as a sponge or a trace fossil. [4] [5]
The Burgess Shale is a fossil-bearing deposit exposed in the Canadian Rockies of British Columbia, Canada. It is famous for the exceptional preservation of the soft parts of its fossils. At 508 million years old, it is one of the earliest fossil beds containing soft-part imprints.
Hexactinellid sponges are sponges with a skeleton made of four- and/or six-pointed siliceous spicules, often referred to as glass sponges. They are usually classified along with other sponges in the phylum Porifera, but some researchers consider them sufficiently distinct to deserve their own phylum, Symplasma. Some experts believe glass sponges are the longest-lived animals on earth; these scientists tentatively estimate a maximum age of up to 15,000 years.
Charles Doolittle Walcott was an American paleontologist, administrator of the Smithsonian Institution from 1907 to 1927, and director of the United States Geological Survey. He is famous for his discovery in 1909 of well-preserved fossils, including some of the oldest soft-part imprints, in the Burgess Shale of British Columbia, Canada.
Vauxia is an extinct genus of demosponge that had a distinctive branching mode of growth. Each branch consisted of a network of strands. Vauxia also had a skeleton of spongin common to modern day sponges. Much like Choia and other sponges, Vauxia fed by extracting nutrients from the water.
The Chancelloriids are an extinct family of superficially sponge-like animals common in sediments from the Early Cambrian to the early Late Cambrian. Many of these fossils consists only of spines and other fragments, and it is not certain that they belong to the same type of organism. Other specimens appear to be more complete and to represent sessile, radially symmetrical hollow bag-like organisms with a soft skin armored with star-shaped calcareous sclerites from which radiate sharp spines.
Chancelloria is a genus of early animals known from the Middle Cambrian Burgess Shale, the Comley limestone, the Wheeler Shale, the Bright Angel Shale and elsewhere. It is named after Chancellor Peak. It was first described in 1920 by Charles Doolittle Walcott, who regarded them as one of the most primitive groups of sponges. However, they are currently thought to be member of the group Chancelloriidae. 178 specimens of Chancelloria are known from the Greater Phyllopod bed, where they comprise 0.34% of the community.
Capsospongia, formerly known as Corralia or Corralio, is a middle Cambrian sponge genus known from 3 specimens in the Burgess shale. Its type and only species is Capsospongia undulata. It has a narrow base, and consists of bulging rings which get wider further up the sponge, resulting in a conical shape. Its open top was presumably used to expel water that had passed through the sponge cells and been filtered for nutrients.
The Burgess Shale, a series of fossil beds in the Canadian Rockies, was first noticed in 1886 by Richard McConnell of the Geological Survey of Canada (GSC). His and subsequent finds, all from the Mount Stephen area, came to the attention of palaeontologist Charles Doolittle Walcott, who in 1907 found time to reconnoitre the area. He opened a quarry in 1910 and in a series of field trips brought back 65,000 specimens, which he identified as Middle Cambrian in age. Due to the quantity of fossils and the pressures of his other duties at the Smithsonian Institution, Walcott was only able to publish a series of "preliminary" papers, in which he classified the fossils within taxa that were already established. In a series of visits beginning in 1924, Harvard University professor Percy Raymond collected further fossils from Walcott's quarry and higher up on Fossil Ridge, where slightly different fossils were preserved.
Hazelia is a genus of spicular Cambrian demosponge known from the Burgess Shale, the Marjum formation of Utah, and possibly Chengjiang. It was described by Charles Walcott in 1920.
Takakkawia is a genus of sponge in the order Protomonaxonida and the family Takakkawiidae. It is known from the Middle Cambrian Burgess Shale that reached around 4 cm in height. Its structure comprises four columns of multi-rayed, organic spicules that align to form flanges. The spicules form blade-like structures, ornamented with concentric rings.
The Phyllopod bed, designated by USNM locality number 35k, is the most famous fossil-bearing member of the Burgess Shale fossil Lagerstätte. It was quarried by Charles Walcott from 1911–1917, and was the source of 95% of the fossils he collected during this time; tens of thousands of soft-bodied fossils representing over 150 genera have been recovered from the Phyllopod bed alone.
Cambrorhytium is an enigmatic fossil genus known from the Latham Shale (California), and the Chengjiang (China) and Burgess Shale lagerstätte. 350 specimens of Cambrorhytium are known from the Greater Phyllopod bed, where they comprise 0.7% of the community.
Eiffelia is an extinct genus of sponges known from the Middle Cambrian Burgess Shale as well as several Early Cambrian small shelly fossil deposits. It is named after Eiffel Peak, which was itself named after the Eiffel Tower. It was first described in 1920 by Charles Doolittle Walcott. It belongs in the Hexactinellid stem group. 60 specimens of Eiffelia are known from the Greater Phyllopod bed, where they comprise 0.11% of the community.
Hamptonia is an extinct genus of sea sponge known from the Middle Cambrian Burgess Shale and the Lower Ordovician Fezouata formation. It was first described in 1920 by Charles Doolittle Walcott. 48 specimens of Hamptonia are known from the Greater Phyllopod bed, where they comprise < 0.1% of the community.
Leptomitus is a genus of demosponge known from the Middle Cambrian Burgess Shale. Its name is derived from the Greek lept ("slender") and mitos ("thread"), referring to the overall shape of the sponge. 138 specimens of Leptomitus are known from the Greater Phyllopod bed, where they comprise 0.26% of the community.
Pirania is an extinct genus of sea sponge known from the Middle Cambrian Burgess Shale and the Ordovician Fezouata formation. It is named after Mount St. Piran, a mountain situated in the Bow River Valley in Banff National Park, Alberta. It was first described in 1920 by Charles Doolittle Walcott. 198 specimens of Pirania are known from the Greater Phyllopod bed, where they comprise 0.38% of the community.
Wapkia is an extinct genus of sea sponge with radial sclerites, known from the Middle Cambrian Burgess Shale. It was first described in 1920 by Charles Doolittle Walcott. 32 specimens of Wapkia are known from the Greater Phyllopod bed, where they comprise 0.06% of the community.
The Mount Stephen trilobite beds are a series of fossil strata on Mount Stephen, British Columbia that contain exceptionally preserved fossil material. Part of the same stratigraphic unit as the Burgess Shale deposit, many non-mineralized parts are preserved; in addition, a high density of trilobite fossils is present.
Protomonaxonida is an extinct order of sea sponges. It is a paraphyletic group gathering the most ancient species from the Burgess Shale to modern sponges.
Amphidiscosida is an order of hexactinellids. The Amphidiscosida are commonly regarded as the only living sponges in the subclass Amphidiscophora.