Burmaculex

Burmaculex; Temporal range: Cretaceous, 95 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Diptera
Family:	Culicidae
Genus:	† Burmaculex ; Borkent & Grimaldi, 2004
Species:	†B. antiquus
Binomial name
	†Burmaculex antiquus; Borkent & Grimaldi, 2004
Other species
	B. edwardsi;

Last updated October 05, 2025

Burmaculex is an extinct monotypic genus of mosquito found fossilised in Burmese amber dating from the Cretaceous period, believed to date from 95 million years ago.^[1] The genus and species were described in 2004 by Art Borkent and David A. Grimaldi.^[2]

Cladogram after Azar et al. (2023),^[3] although Libanoculex is later confirmed as member of Chaoboridae and not a mosquito.^[4]

Corethrellidae

Chaoboridae

† Libanoculex intermedius (Lebanese amber)

	†Burmaculex antiquusBorkent & Grimaldi, 2004 (Burmese amber)^[2]

	Crown Culicidae

References

↑ Quentin D. Wheeler (6 December 2012). "New to nature No 95: Culiseta lemniscata". The Guardian . Retrieved 23 July 2022.
1 2 Borkent, Art; Grimaldi, David A. (1 September 2004). "The Earliest Fossil Mosquito (Diptera: Culicidae), in Mid-Cretaceous Burmese Amber". Annals of the Entomological Society of America . 97 (5): 882–888. doi: 10.1603/0013-8746(2004)097[0882:TEFMDC]2.0.CO;2 . S2CID 85738337.
↑ Azar, Dany; Nel, André; Huang, Diying; Engel, Michael S. (December 2023). "The earliest fossil mosquito". Current Biology. 33 (23): 5240–5246.e2. doi: 10.1016/j.cub.2023.10.047 .
↑ Harbach, Ralph E. (2024-03-12). "Libanoculex intermedius is not a mosquito (Diptera: Culicidae): It is a chaoborid (Chaoboridae)". Zootaxa. 5424 (1): 139–144. doi:10.11646/zootaxa.5424.1.9. ISSN 1175-5334.

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[The_Guardian_-_6_December_2012_-_New_to_nature_No_95:_Culiseta_lemniscata-1] Quentin D. Wheeler (6 December 2012). "New to nature No 95: Culiseta lemniscata". The Guardian . Retrieved 23 July 2022.

[Borkent2004-2] 1 2 Borkent, Art; Grimaldi, David A. (1 September 2004). "The Earliest Fossil Mosquito (Diptera: Culicidae), in Mid-Cretaceous Burmese Amber". Annals of the Entomological Society of America . 97 (5): 882–888. doi: 10.1603/0013-8746(2004)097[0882:TEFMDC]2.0.CO;2 . S2CID 85738337.

[Azar2023-3] Azar, Dany; Nel, André; Huang, Diying; Engel, Michael S. (December 2023). "The earliest fossil mosquito". Current Biology. 33 (23): 5240–5246.e2. doi: 10.1016/j.cub.2023.10.047 .

[4] Harbach, Ralph E. (2024-03-12). "Libanoculex intermedius is not a mosquito (Diptera: Culicidae): It is a chaoborid (Chaoboridae)". Zootaxa. 5424 (1): 139–144. doi:10.11646/zootaxa.5424.1.9. ISSN 1175-5334.

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