Burmaculex

Burmaculex; Temporal range: Cretaceous, 95 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Diptera
Family:	Culicidae
Genus:	† Burmaculex ; Borkent & Grimaldi, 2004
Type species
	Burmaculex antiquus; Borkent & Grimaldi, 2004
Species
	Burmaculex antiquus Borkent & Grimaldi, 2004; Burmaculex edwardsi Szadziewski et al., 2024; Burmaculex harbachi Szadziewski et al., 2024; Burmaculex porczynskii Szadziewski et al., 2024 ;

Last updated December 06, 2025

Burmaculex is an extinct genus of mosquito found fossilised in Burmese amber dating from the Cretaceous period, believed to date from 95 million years ago.^[1] The genus and species were described in 2004 by Art Borkent and David A. Grimaldi.^[2]

Cladogram after Azar et al. (2023),^[3] although Libanoculex is later confirmed as member of Chaoboridae and not a mosquito.^[4]

Corethrellidae

Chaoboridae

† Libanoculex intermedius (Lebanese amber)

	†Burmaculex antiquusBorkent & Grimaldi, 2004 (Burmese amber)^[2]

	Crown Culicidae

References

↑ Quentin D. Wheeler (6 December 2012). "New to nature No 95: Culiseta lemniscata". The Guardian . Retrieved 23 July 2022.
1 2 Borkent, Art; Grimaldi, David A. (1 September 2004). "The Earliest Fossil Mosquito (Diptera: Culicidae), in Mid-Cretaceous Burmese Amber". Annals of the Entomological Society of America . 97 (5): 882–888. doi: 10.1603/0013-8746(2004)097[0882:TEFMDC]2.0.CO;2 . S2CID 85738337.
↑ Azar, Dany; Nel, André; Huang, Diying; Engel, Michael S. (December 2023). "The earliest fossil mosquito". Current Biology. 33 (23): 5240–5246.e2. doi: 10.1016/j.cub.2023.10.047 .
↑ Harbach, Ralph E. (2024-03-12). "Libanoculex intermedius is not a mosquito (Diptera: Culicidae): It is a chaoborid (Chaoboridae)". Zootaxa. 5424 (1): 139–144. doi:10.11646/zootaxa.5424.1.9. ISSN 1175-5334.

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[The_Guardian_-_6_December_2012_-_New_to_nature_No_95:_Culiseta_lemniscata-1] Quentin D. Wheeler (6 December 2012). "New to nature No 95: Culiseta lemniscata". The Guardian . Retrieved 23 July 2022.

[Borkent2004-2] 1 2 Borkent, Art; Grimaldi, David A. (1 September 2004). "The Earliest Fossil Mosquito (Diptera: Culicidae), in Mid-Cretaceous Burmese Amber". Annals of the Entomological Society of America . 97 (5): 882–888. doi: 10.1603/0013-8746(2004)097[0882:TEFMDC]2.0.CO;2 . S2CID 85738337.

[Azar2023-3] Azar, Dany; Nel, André; Huang, Diying; Engel, Michael S. (December 2023). "The earliest fossil mosquito". Current Biology. 33 (23): 5240–5246.e2. doi: 10.1016/j.cub.2023.10.047 .

[4] Harbach, Ralph E. (2024-03-12). "Libanoculex intermedius is not a mosquito (Diptera: Culicidae): It is a chaoborid (Chaoboridae)". Zootaxa. 5424 (1): 139–144. doi:10.11646/zootaxa.5424.1.9. ISSN 1175-5334.

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