Capistrano Formation

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Capistrano Formation
Stratigraphic range: Miocene to Pliocene 6.6–4.9  Ma
O
S
D
C
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Pg
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Type Formation
Sub-unitsOso Member, unnamed siltstone member
Underlies Niguel Formation
Overlies Monterey Formation
Thickness850 m
Lithology
Primary Siltstone, Arkose
Other Breccia, Mudstone
Location
Region Orange County, California
Country United States
Type section
Named for San Juan Capistrano
Named byAlfred Oswald Woodford
Year defined1925
Oso Member.jpg
Outcrops of the Oso Member, with the type locality of Titanotaria highlighted

The Capistrano Formation is a geologic formation in coastal southern Orange County, California. It preserves fossils dating back to the late Miocene to early Pliocene, with the Oso Member representing a near-shore environment. Fifty-nine species and varieties of foraminifera are recognized from the Capistrano Formation [1] alongside a diverse array of marine mammals including up to five species of walrus.

Contents

Geography

The Capistrano Formation, named for the town of San Juan Capistrano, is located in southern California, specifically the northern extent of the Peninsular Ranges, which stretch from the Los Angeles Basin to Baja California. [2] It crops out along the coast from Dana Point to San Clemente, and inland for seven miles. [1]

Geology and Stratigraphy

The Capistrano Formation is a heterogenous marine formation that can be differentiated into two distinct but adjacent units. One of these units is the Oso Member, which is composed of arkosic sandstone and preserves a nearshore environment. This unit shows outcrops across Orange County, which are recognizable as medium to coarse grained, white to tan rock. The second member primarily consists of siltstone, preserves an environment that would have been located further out at sea in deeper waters and has not yet been named. At the southwestern border of the Oso Member the two units connect.

Depending on the locality, the Oso Member of the Capistrano Formation either conformably overlies the Puente Formation (specifically Soquel Member), while in other areas it unconformably overlies the Monterey Formation and is succeeded by the Niguel Formation. The Oso Member has been dated to the early late Hemphillian (6.6 to 5.8 Ma) based on the presence of Dinohippus interpolatus and absence of older or younger taxa, but the formation as a whole extends into the Pliocene, rendering the Oso Member coeval with only the lower strata of the unnamed siltstone member. Foraminifers and microfossils have also previously been used to date the formation, which indicate an early Pliocene age (5.6 to 4.9 Ma) for its upper boundary. [3]

Paleoenvironment

The two units of the Capistrano Formation preserve environments of difference distance from the shore. The Oso Member preserves a near-shore environment, thought to be submarine delta deposits situated in a shallow embayment of the Pacific Ocean. The presence of marlin fossils could indicate that parts of the formation deposited at bathyal depths, between 200 and 2.000 meters deep, [3] and Fierstine argues that the preservation of the material suggests that it was not subject to extensive postmortem transportation. He also argues that the presence of this fish suggests warm water temperatures during the Miocene, with Marlins typically preferring average surface temperatures of 24 °C. [4] Barboza and colleagues meanwhile argue that this alone is not sufficient evidence for depth, citing the preservation of marlin fossils in much shallower waters including one specimen found in deposits of the supralittoral zone. [2] The proximity to the shore is apparent due to the presence of terrestrial fauna.

Paleofauna

Chondrichthyes

NameSpeciesMemberMaterialNotesImage
Cosmopolitodus C. hastalis [2] Oso Memberalso known as the broad-toothed mako
Cosmopolitodus hastalis tooth Cosmopolitodus hastalis (fossil shark tooth) (Tertiary; probably from the Chesapeake Bay region, USA) 3.jpg
Cosmopolitodus hastalis tooth
Myliobatis M. sp. [2] Oso Membera species of ray
Otodus O. megalodon [5] Oso Member

Siltstone Member

teeth
Megalodon restoration Megalodon restoration.png
Megalodon restoration

Osteichthyes

NameSpeciesMemberMaterialNotesImage
Acipenseridae indet. [6] Oso Membera type of sturgeon
Makaira M. nigricans [4] Oso Membera nearly complete skullan early specimen of the extant blue marlin
Blue marlin Blue marlin (Duane Raver).png
Blue marlin
Oncorhynchus O. rastrosus [7] Oso Memberthe sabertoothed salmon
Oncorhynchus rastrosus Oncorhynchus rastrosus cropped.png
Oncorhynchus rastrosus

Reptilia

NameSpeciesMemberMaterialNotesImage
Crocodilia indet. [2] Oso MemberA tooth and an osteoderm
Dermochelyidae indet. [2] Oso Member
Testudinidae indet. [2] Oso Member

Aves

NameSpeciesMemberMaterialNotesImage
Mancallinae indet. [8] [9] Oso Membera type of flightless auk
Uria U. sp. [8]

Mammalia

Afrotheria

NameSpeciesMemberMaterialNotesImage
Desmostylus D. sp. [2] Oso Member
Desmostylus Desmostylus ROM.jpg
Desmostylus
Hydrodamalis H. cuestae [2] Oso Membera close relative of Steller's Sea Cow
Proboscidea indet. [2] Oso Member

Artiodactyla

NameSpeciesMemberMaterialNotesImage
Antilocapridae indet. [2] Oso Member
cf. Balaenoptera [2] cf. Balaenoptera sp.Oso Member
Camelidae indet. [2] Oso Member
Delphinidae indet. [2] Oso Member
Herpetocetus [10] Oso Membera cetotheriid whale
Herpetocetus bramblei Herpetocetus bramblei.png
Herpetocetus bramblei
Parapontoporia P. pacifica [11] Oso Membera skull with associated petrotympanica relative of the Chinese river dolphin
Parapontoporia sternbergi (left), a related species Fossil whales.jpg
Parapontoporia sternbergi (left), a related species
Physeteroidea Species A [2] Oso Member
Physeteroidea Species B [2] Oso Member
Tayassuidae indet. [2] Oso Member

Carnivora

NameSpeciesMemberMaterialNotesImage
Borophagus [2] Oso Membera large genus of canid
Borophagus skull Osteoborus cyanoides.jpg
Borophagus skull
Mustelidae indet. [2] Oso Member
Gomphotaria G. pugnax [8] Oso Membera species of walrus
Gomphotaria pugnax life restoration Gomphotaria pugnax life restoration.jpg
Gomphotaria pugnax life restoration
Osodobenus O. eodon [12] Oso Memberthree skulls representing an adult male, adult female and juvenile specimenthe first known walrus with tusks
Pontolis cf. P. magnus

P. kohnoi [12]

Oso Membera giant early walrus
Pontolis magnus mandible Pontolis magnus.jpg
Pontolis magnus mandible
Thalassoleon T. mexicanus [3] Siltstone Membermultiple skulls and postcranial materiala basal eared seal
Thalassoleon mexicanus skull Thalassoleon mexicanus skull 2.jpg
Thalassoleon mexicanus skull
Titanotaria T. orangensis [13] Oso Memberskull and postcranial material of multiple individualsa basal walrus
Lateral view of Titanotaria orangensis Titanotaria lateral.jpg
Lateral view of Titanotaria orangensis

Lagomorpha

NameSpeciesMemberMaterialNotesImage
Leporidae indet. [2] Oso Member

Perissodactyla

NameSpeciesMemberMaterialNotesImage
Dinohippus [2] D. interpolatusOso Memberisolated teeth
Rhinocerotidae indet. [2] Oso Member

See also

Related Research Articles

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References

  1. 1 2 White, W.R. (1956). "Pliocene and Miocene Foraminifera from the Capistrano Formation, Orange County, California". Journal of Paleontology. 30 (2): 237–260. JSTOR   1300262.
  2. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 Barboza, M.M.; Parham, J.F.; Santos, G.; Kussman, B.N.; Velez-Juarbe, J. (2017). "The age of the Oso Member, Capistrano Formation, and a review of fossil crocodylians from California". PaleoBios. 34. doi: 10.5070/P9341033797 .
  3. 1 2 3 Deméré, T.A.; Berta, A. (2005). "New skeletal material of Thalassoleon (Otariidae Pinnipedia) from the late Miocene-early Pliocene (Hemphillian) of California". Bulletin of the Florida Museum of Natural History. 45 (4): 379–411.
  4. 1 2 Fierstine, H. L. (2008). "A Fossil Skull of the Extant Blue Marlin (Makaira nigricans Lacepède, 1802) from the Late Miocene of Orange County, California". Bulletin, Southern California Academy of Sciences. 107 (2): 45–56. doi:10.3160/0038-3872(2008)107[45:AFSOTE]2.0.CO;2. S2CID   26337210.
  5. Boessenecker, R.W.; Ehret, D.J.; Long, D.J.; Churchill, M.; Martin, E.; Boessenecker, S.J. (2019). "The Early Pliocene extinction of the mega-toothed shark Otodus megalodon: a view from the eastern North Pacific". PeerJ. 7: e6088. doi: 10.7717/peerj.6088 . PMC   6377595 . PMID   30783558. S2CID   67750037.
  6. Hilton, E.J.; Grande, L. (2017). "Review of the fossil record of sturgeons, family Acipenseridae (Actinopterygii: Acipenseriformes), from North America". Journal of Paleontology. 80 (4): 672–683. doi:10.1666/0022-3360(2006)80[672:ROTFRO]2.0.CO;2. S2CID   53583501.
  7. Sankey, J.; Biewer, J.; Basuga, J.; Palacios, F.; Wagner, H.; Garber, D. (2016). "The giant, spike-toothed salmon, Oncorhynchus rastrosus and the "Proto-Tuolumne River" (early Pliocene) of Central California" (PDF). PaleoBios. 33. doi:10.5070/P9331033123.
  8. 1 2 3 Smith, N. A.; Boessenecker, R.; Long, D.; Powell, C. "A new marine vertebrate assemblage from the Wilson Grove Formation at Bloomfield Quarry (late Miocene), Sonoma County, California". Journal of Vertebrate Paleontology, Programs and Abstracts 2017, 37:195.
  9. Morales, D.; Kloess, P.A.; Parham, J.F. (2015). "On the abundance of flightless auks from the middle Miocene to early Pliocene" (PDF). PaleoBios. 23 (1).
  10. Tanaka, Y.; Furusawa, H.; Barnes, L. G. (2018). "Fossil Herpetocetine Baleen Whales (Cetacea, Mysticeti, Cetotheriidae) from the Lower Pliocene Horokaoshirarika Formation at Numata, Hokkaido, Northern Japan". Paleontological Research. 22 (4): 295–306. doi:10.2517/2017pr025. S2CID   134686678.
  11. Boessenecker, R.W.; Poust, A.W. (2015). "Freshwater occurrence of the extinct dolphin Parapontoporia (Cetacea: Lipotidae) from the upper Pliocene nonmarine Tulare Formation of California". Palaeontology. 58 (3): 489–496. doi: 10.1111/pala.12153 . S2CID   128980362.
  12. 1 2 Biewer, Jacob N.; Velez-Juarbe, Jorge; Parham, James F. (2020-12-01). "Insights on the Dental Evolution of Walruses Based on New Fossil Specimens from California". Journal of Vertebrate Paleontology. 40 (5): e1833896. doi:10.1080/02724634.2020.1833896. ISSN   0272-4634. S2CID   228814992.
  13. Magallanes, I.; Parham, J.F.; Santos, G.; Velez-Juarbe, J. (2018). "A new tuskless walrus from the Miocene of Orange County, California, with comments on the diversity and taxonomy of odobenids". PeerJ. 6: e5708. doi: 10.7717/peerj.5708 . PMC   6188011 . PMID   30345169. S2CID   53020427.