Capsospongia Temporal range: Middle Cambrian, | |
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Artist's restoration | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Porifera |
Class: | Demospongiae |
Family: | † Anthaspidellidae |
Genus: | † Capsospongia Rigby 1986 |
Species: | †C. undulata |
Binomial name | |
†Capsospongia undulata Walcott 1920 | |
Capsospongia, formerly known as Corralia or Corralio, is a middle Cambrian sponge genus known from 3 specimens in the Burgess shale. [1] Its type and only species is Capsospongia undulata. It has a narrow base, and consists of bulging rings which get wider further up the sponge, resulting in a conical shape. Its open top was presumably used to expel water that had passed through the sponge cells and been filtered for nutrients.
Like most sponges, Capsospoingia had a spicular skeleton; long spicules parallel to the growth direction formed columns which were connected by shorter lateral spicules.
Capsospongia undulata was named in 1920 by Charles Walcott as Corralia undulata. [2] However, the name was preoccupied by Corralia Roewer, 1913, a member of Opiliones. In 1955, de Laubenfels renamed the genus Corralio, adopting an incorrect spelling of Corralia Walcott had used. [3] [4] In 1986 Keith Rigby established the new genus Capsospongia for it. [5] In 2004, he and Desmond Collins described a third specimen. [5]
C. undulata intersects with the complicated taxonomic history of the anomalocarids. In 1911, Walcott had named two taxa, Peytoia and Laggania, which he interpreted as a jellyfish and a sea cucumber respectively. [6] In 1978, Simon Conway Morris recognized that the mouthparts of Laggania closely resembled Peytoia, but erroneously concluded that this was because Laggania was a composite fossil of a Peytoia and another organism, which he concluded was a sponge and suggested was probably a specimen of C. undulata. [4] However, it was subsequently determined that Laggania and Peytoia were partial specimens of a larger animal, a radiodont, which now bears the name Peytoia. [7] [8]
Anomalocaris is an extinct genus of radiodont, an order of early-diverging stem-group arthropods.
Peytoia is a genus of hurdiid radiodont, an early diverging order of stem-group arthropods, that lived in the Cambrian period, containing two species, Peytoia nathorsti from the Miaolingian of Canada and Peytoia infercambriensis from Poland, dating to Cambrian Stage 3. Its two frontal appendages had long bristle-like spines, it had no fan tail, and its short stalked eyes were behind its large head.
Choia is a genus of extinct demosponge ranging from the Cambrian until the Lower Ordovician periods. Fossils of Choia have been found in the Burgess Shale in British Columbia; the Maotianshan shales of China; the Wheeler Shale in Utah; and the Lower Ordovician Fezouata formation. It was first described in 1920 by Charles Doolittle Walcott.
Peytoia infercambriensis is a species of hurdiid radiodont in the genus Peytoia.
The Chancelloriids are an extinct family of superficially sponge-like animals common in sediments from the Early Cambrian to the early Late Cambrian. Many of these fossils consists only of spines and other fragments, and it is not certain that they belong to the same type of organism. Other specimens appear to be more complete and to represent sessile, radially symmetrical hollow bag-like organisms with a soft skin armored with star-shaped calcareous sclerites from which radiate sharp spines.
Chancelloria is a genus of early animals known from the Middle Cambrian Burgess Shale, the Comley limestone, the Wheeler Shale, the Bright Angel Shale and elsewhere. It is named after Chancellor Peak. It was first described in 1920 by Charles Doolittle Walcott, who regarded them as one of the most primitive groups of sponges. However, they are currently thought to be member of the group Chancelloriidae. 178 specimens of Chancelloria are known from the Greater Phyllopod bed, where they comprise 0.34% of the community.
Radiodonta is an extinct order of stem-group arthropods that was successful worldwide during the Cambrian period. They may be referred to as radiodonts, radiodontans, radiodontids, anomalocarids, or anomalocaridids, although the last two originally refer to the family Anomalocarididae, which previously included all species of this order but is now restricted to only a few species. Radiodonts are distinguished by their distinctive frontal appendages, which are morphologically diverse and used for a variety of functions. Radiodonts included the earliest large predators known, but they also included sediment sifters and filter feeders. Some of the most famous species of radiodonts are the Cambrian taxa Anomalocaris canadensis, Hurdia victoria, Peytoia nathorsti, Titanokorys gainessii, Cambroraster falcatus and Amplectobelua symbrachiata, the Ordovician Aegirocassis benmoulai and the Devonian Schinderhannes bartelsi.
Hurdia is an extinct genus of hurdiid radiodont that lived 505 million years ago during the Cambrian Period. Fossils have been found in North America, China and the Czech Republic.
The Burgess Shale, a series of fossil beds in the Canadian Rockies, was first noticed in 1886 by Richard McConnell of the Geological Survey of Canada (GSC). His and subsequent finds, all from the Mount Stephen area, came to the attention of palaeontologist Charles Doolittle Walcott, who in 1907 found time to reconnoitre the area. He opened a quarry in 1910 and in a series of field trips brought back 65,000 specimens, which he identified as Middle Cambrian in age. Due to the quantity of fossils and the pressures of his other duties at the Smithsonian Institution, Walcott was only able to publish a series of "preliminary" papers, in which he classified the fossils within taxa that were already established. In a series of visits beginning in 1924, Harvard University professor Percy Raymond collected further fossils from Walcott's quarry and higher up on Fossil Ridge, where slightly different fossils were preserved.
Takakkawia is a genus of sponge in the order Protomonaxonida and the family Takakkawiidae. It is known from the Middle Cambrian Burgess Shale that reached around 4 cm in height. Its structure comprises four columns of multi-rayed, organic spicules that align to form flanges. The spicules form blade-like structures, ornamented with concentric rings.
Cambrorhytium is an enigmatic fossil genus known from the Latham Shale (California), and the Chengjiang (China) and Burgess Shale lagerstätte. 350 specimens of Cambrorhytium are known from the Greater Phyllopod bed, where they comprise 0.7% of the community.
Eiffelia is an extinct genus of sponges known from the Middle Cambrian Burgess Shale as well as several Early Cambrian small shelly fossil deposits. It is named after Eiffel Peak, which was itself named after the Eiffel Tower. It was first described in 1920 by Charles Doolittle Walcott. It belongs in the Hexactinellid stem group. 60 specimens of Eiffelia are known from the Greater Phyllopod bed, where they comprise 0.11% of the community.
Halichondrites, sometimes mis-spelt Halicondrites is an extinct genus of sea sponge known from the Middle Cambrian Burgess Shale. 7 specimens of Halichondrites are known from the Greater Phyllopod bed, where they comprise < 0.1% of the community.
Hamptonia is an extinct genus of sea sponge known from the Middle Cambrian Burgess Shale and the Lower Ordovician Fezouata formation. It was first described in 1920 by Charles Doolittle Walcott. 48 specimens of Hamptonia are known from the Greater Phyllopod bed, where they comprise < 0.1% of the community.
Louisella is a genus of worm known from the Middle Cambrian Burgess Shale. It was originally described by Charles Walcott in 1911 as a holothurian echinoderm, and represents a senior synonym of Miskoia, which was originally described as an annelid. 48 specimens of Louisella are known from the Greater Phyllopod bed, where they comprise < 0.1% of the community. It has been stated to have palaeoscolecid-like sclerites, though this is not in fact the case.
Leptomitus is a genus of demosponge known from the Middle Cambrian Burgess Shale. Its name is derived from the Greek lept ("slender") and mitos ("thread"), referring to the overall shape of the sponge. 138 specimens of Leptomitus are known from the Greater Phyllopod bed, where they comprise 0.26% of the community.
Wapkia is an extinct genus of sea sponge with radial sclerites, known from the Middle Cambrian Burgess Shale. It was first described in 1920 by Charles Doolittle Walcott. 32 specimens of Wapkia are known from the Greater Phyllopod bed, where they comprise 0.06% of the community.
The Mount Stephen trilobite beds are a series of fossil strata on Mount Stephen, British Columbia that contain exceptionally preserved fossil material. Part of the same stratigraphic unit as the Burgess Shale deposit, many non-mineralized parts are preserved; in addition, a high density of trilobite fossils is present.
Lecythioscopa is a genus of probable archaeopriapulid known from two specimens from the Walcott Quarry from the Middle Cambrian Burgess Shale.
Protomonaxonida is an extinct order of sea sponges. It is a paraphyletic group gathering the most ancient species from the Burgess Shale to modern sponges.