Cistus heterophyllus | |
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Cistus heterophyllus subsp. carthaginensis, Sierra Minera de Cartagena, Spain | |
Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Angiosperms |
Clade: | Eudicots |
Clade: | Rosids |
Order: | Malvales |
Family: | Cistaceae |
Genus: | Cistus |
Species: | C. heterophyllus |
Binomial name | |
Cistus heterophyllus Desf. [1] | |
Cistus heterophyllus is a shrubby species of flowering plant in the family Cistaceae.
Cistus heterophyllus grows up to 1 m (3 ft) tall, forming an erect, much-branched shrub. Its leaves are elliptical to lanceolate in shape, usually 5–20 mm (0.2–0.8 in) long, the upper surfaces being dark green with stellate and simple hairs, and the lower surfaces whitish with a coating of short hairs. The leaf margins are slightly turned under (revolute) and the veins are much more obvious on the underside. The leaves are of two kinds: the upper are without stalks (petioles), the lower have short stalks. The flowers are arranged in cymes of one to five individual flowers, each with five purplish-pink petals, usually with a yellow spot at the base. Like the leaves, the five sepals have stellate hairs, plus some longer simple hairs. The fruit capsule is about 9 mm (0.35 in) high containing angular brownish seeds. [2] [3]
The two described subspecies differ in the distribution of hairs. In C. h. subsp. heterophyllus, the young stems have both many stellate hairs and many longer simple hairs, and the leaves have scattered long simple hairs. In C. h. subsp. carthaginensis, the young stems and leaves have fewer simple hairs. The outer two sepals of C. h. subsp. heterophyllus average about 10 mm (0.39 in) long by 9 mm (0.35 in) wide, whereas those of C. h. subsp. carthaginensis are smaller, averaging about 8.5 mm (0.33 in) long by 6 mm (0.24 in) wide. The petals of C. h. subsp. heterophyllus are about 20–30 mm (0.8–1.2 in) long by 20–25 mm (0.8–1.0 in) wide, making the flower diameter typically 40–50 mm (1.6–2.0 in). Although there is considerable variation and the size ranges overlap, the petals of C. h. subsp. carthaginensis may be as small as 12 mm (0.47 in) long and 10 mm (0.39 in) wide, making the flower diameter typically 30–40 mm (1.2–1.6 in). [4]
Cistus heterophyllus was first described by René Louiche Desfontaines in 1798. [1] The specific epithet heterophyllus means "with leaves of different shapes", [5] the upper being unstalked, the lower shortly stalked.
Two subspecies are accepted: [1] [6]
The two subspecies have been artificially crossed forming a hybrid subspecies called C. h. nothosubsp. marzoi. [4]
A 2011 molecular phylogenetic study placed C. heterophyllus in the purple and pink flowered clade (PPC) of Cistus species, in a subclade with C. albidus and C. creticus . [7]
Species-level cladogram of Cistus species. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Species-level cladogram of Cistus species, based on plastid and nuclear DNA sequences. [7] [8] [9] [10] |
Cistus heterophyllus has a disjoint distribution. C. h. subsp. heterophyllus is native to western North Africa, along the coastal Mediterranean region from the Spanish island of Peñón de Alhucemas and Targuist in Morocco to Algiers. [4] C. h. subsp. carthaginensis is found only in two locations in mainland Spain: near La Pobla de Vallbona in Valencia and in the Calblanque Regional Park in Murcia. [6]
Cistus heterophyllus subsp. carthaginensis is listed as critically endangered (CR) in the IUCN Red List. [11] Fewer than 30 individuals were known in 2011 (only one at the Valencian site) and some appear to be hybrids with Cistus albidus . [6] The subspecies is threatened by fires and human disturbance. Seeds have been stored in seedbanks, and attempts are being made to increase plants in cultivation, both from seed and in vitro. [11]
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Cistus laurifolius, commonly called laurel-leaf cistus, laurel-leaved cistus or laurel-leaved rock rose, is a species of highly branched flowering evergreen shrub native to some areas around the Mediterranean.
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Cistus albidus, the grey-leaved cistus, is a shrubby species of flowering plant in the family Cistaceae, with pink to purple flowers, native to south-western Europe and western north Africa.
Cistus asper is a shrubby species of flowering plant in the family Cistaceae with purple-pink flowers. It was first described in 2005 and is endemic to El Hierro in the Canary Islands.
Cistus chinamadensis is a shrubby species of flowering plant in the family Cistaceae, with purple-pink flowers, first described in 1991. It is endemic to the Canary Islands, where three subspecies occur on three separate islands. The species has been assessed as endangered in the IUCN Red List, being known only from small separated areas and facing a variety of threats.
Cistus clusii is a shrubby species of flowering plant in the family Cistaceae, with white flowers, native to south west and south central Europe and north Africa. It has been wrongly called Cistus libanotis by many authors.
Cistus crispus is a shrubby species of flowering plant in the family Cistaceae, with pink to purple flowers, native to south-western Europe and western north Africa.
Cistus horrens is a shrubby species of flowering plant in the family Cistaceae, with purple to pink flowers. It is endemic to Gran Canaria in the Canary Islands. First described as a species in 2004, it was previously identified as Cistus symphytifolius, which it resembles.
Cistus inflatus is a shrubby species of flowering plant in the family Cistaceae, often known as Cistus psilosepalus, although this name is a synonym of the hybrid Cistus × laxus. It has white flowers.
Cistus libanotis is a shrubby species of flowering plant in the family Cistaceae, with white flowers. It has been confused with Cistus clusii, which it resembles, resulting in some uncertainty in its distribution. It is endemic to the Iberian Peninsula.
Cistus munbyi is a shrubby species of flowering plant in the family Cistaceae, with white flowers. Related to and resembling Cistus clusii, it is native to Morocco and Algeria in western north Africa.
Cistus ocreatus is a shrubby species of flowering plant in the family Cistaceae, with purple-pink flowers. It is sometimes treated as synonymous with Cistus symphytifolius or as its subspecies C. symphytifolius subsp. leucophyllus. Its name is sometimes spelt Cistus ochreatus. It is endemic to Gran Canaria in the Canary Islands.
Cistus osbeckiifolius is a shrubby species of flowering plant in the family Cistaceae, with pink to purple flowers.
Cistus palmensis is a shrubby species of flowering plant in the family Cistaceae.
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