Trilobites are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last trilobites disappeared in the mass extinction at the end of the Permian about 251.9 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described.
Phacops is a genus of trilobites in the order Phacopida, family Phacopidae, that lived in Europe, northwestern Africa, North and South America and China from the Late Ordovician until the very end of the Devonian, with a broader time range described from the Late Ordovician. It was a rounded animal, with a globose head and large eyes, and probably fed on detritus. Phacops is often found rolled up ("volvation"), a biological defense mechanism that is widespread among smaller trilobites but further perfected in this genus.
Floresta is a town and municipality in Boyacá Department, Colombia, part of the Tundama Province, a subregion of Boyacá. Floresta is situated on the Altiplano Cundiboyacense in the Eastern Ranges of the Colombian Andes. It borders Santa Rosa de Viterbo, Cerinza and Betéitiva in the north, in the east Busbanzá and Corrales in the west Santa Rosa de Viterbo and Nobsa in the south.
Holoptychius is an extinct genus of porolepiform lobe-finned fish from the Middle Devonian to Carboniferous (Mississippian) periods. It is known from fossils worldwide. The genus was first described by Louis Agassiz in 1839.
Greenops is a mid-sized Devonian trilobite of the order Phacopida, subfamily Asteropyginae. They are mainly reported from the mid-Devonian Hamilton Group of upstate New York and southwestern Ontario. A similar-looking trilobite from Morocco is often mis-labelled Greenops. Greenops had schizocroidal eyes, large genal spines and short, sharp spines at the tip of each segment of the pygidium ("tail"). Greenops lived in warm, fairly deep water. In the Hamilton Group of New York, they are found with Eldredgeops, Dipleura and Bellacartwrightia, a trilobite that resembles Greenops but has much larger pygidial spines. In Ontario, they are found in the Widder Formation, which outcrops at Arkona, where they are, by far, the dominant trilobite.
Gravicalymene Shirley, 1936, is a genus of trilobites belonging to the order Phacopida, suborder Calymenina and family Calymenidae. Species included in this genus have previously been allocated to Calymene Brongniart 1822,Flexicalymene Shirley, 1936. and Sthenarocalymene Siveter 1977.
Atrypa is a genus of brachiopod with round to short egg-shaped shells covered with many fine radial ridges. Growth lines form perpendicular to the costae and are spaced approximately 2 to 3 times further apart than the costae.. The pedunculate valve is slightly convex, but oftentimes levels out or becomes slightly concave toward the anterior margin. The brachial valve is highly convex. Neither valve contains an interarea. Atrypa had a large geographic range and occurred from the late Lower Silurian (Telychian) to the early Upper Devonian (Frasnian). Other sources expand the range from the Late Ordovician to Carboniferous, approximately from 449 to 336 Ma. A proposed new species, A. harrisi, was found in the trilobite-rich Floresta Formation in Boyacá, Colombia.
Erbenochile is a genus of spinose phacopid trilobite, of the family Acastidae, found in Lower to Middle Devonian age rocks from Algeria and Morocco. Originally described from an isolated pygidium, the first complete articulated specimen of E. erbeni revealed the presence of extraordinarily tall eyes:
"Straight-sided towers of lenses... with [up to] 18 lenses in a vertical file"
Cyphaspis is a genus of small trilobite that lived from the Late Ordovician to the Late Devonian. Fossils have been found in marine strata in what is now Europe, Africa and North America. Various species had a compact body, and a large, bulbous glabellum. Many species had long spines arranged similarly to closely related genera, such as Otarian, Otarionella, Chamaeleoaspis, and Namuropyge.
Anchiopsis is a genus of trilobite in the order Phacopida, which existed in what is now New York, U.S.A. It was described by Delo in 1935, and the type species is Anchiopsis anchiops, which was originally described as Calymene anchiops by Green in 1832.
Tarijactinoides is a genus of trilobites in the order Phacopida, which existed in what is now Bolivia. It was described by Suárez Soruco in 1971, and the type species is Tarijactinoides jarcasensis.
Coronura is a trilobite in the order Phacopida, that existed during the Middle Devonian in what is now New York, U.S.A. It was described by Hall and Clarke in 1888, and the type species is Coronura aspectans, which was originally described under the genus Asaphus by Conrad in 1841. The species was described from the Onondaga Formation. Fossils of Coronura have also been found in Indiana, and in the Emsian to Givetian Floresta Formation, Altiplano Cundiboyacense, Colombia.
Homalonotus is an extinct genus of trilobites in the order Phacopida. It contains several species, including H. armatus and H. roemeri. It is closely related to other trilobites such as Arduennella and Dipleura..
Platyceras is a genus of extinct Paleozoic sea snails, marine gastropod mollusks in the family Platyceratidae. This genus is known from the Silurian to the Middle Triassic periods and especially abundant in the Devonian and Carboniferous. It is the type genus of the family Platyceratidae.
Marrellomorpha are an extinct group of arthropods known from the Cambrian to the Early Devonian. They lacked mineralised hard parts, so are only known from areas of exceptional preservation, limiting their fossil distribution. The best known member is Marrella, with thousands of specimens found in the Cambrian aged Burgess Shale of Canada. The group is divided up into two major orders, Marrellida and Acercostraca. Marrellida is recognised by the possession of head shields with two or three pairs of elongate spine-like projections, and three pairs of uniramous appendages on the cephalon, while Acercostraca generally have large ovoid carapaces that cover the entire upper half of the body, and five pairs of uniramous cephalic appendages. Both groups have unbranched antennules and a segmented trunk with biramous appendages. Recent research has suggested the previously enigmatic Cambrian trilobite-like arthropods Skania and Primicaris belong to this group. Their phylogenetic position is uncertain, various studies have alternatively placed them in the Arachnomorpha as relatives of Artiopoda, as related to Mandibulata, or as stem group euarthropods.
Cordania is an extinct genus of trilobites that lived from the Early to Middle Devonian.
Viaphacops is a genus of trilobites in the order Phacopida, family Phacopidae, that lived during the Middle Devonian, and is known from North and South America, Asia.
Odontopleura is a genus of spinose odontopleurid trilobite in the family Odontopleuridae, and is the type genus of that family and of Odontopleurida. The various species are found in Upper Ordovician to Middle Devonian marine strata throughout the world. The best studied fossils are of the type species, O. ovata, from the Wenlock-aged Liteň Formation in Loděnice, in Bohemia, Czech Republic, and, southeastern Gotland, of Sweden.
The Floresta Formation is a geological formation of the Altiplano Cundiboyacense in the Eastern Ranges of the Colombian Andes. The sequence of siltstones, shales, coquinas and sandstone beds dates to the Devonian period; Late Emsian, Eifelian and Early Givetian epochs, and has a maximum thickness of 600 metres (2,000 ft). The unit is highly fossiliferous; brachiopods, bryozoans, gastropods, trilobites, corals and bivalves have been found in the Floresta Formation. Some fragments of Placoderm fish fossils were found in the Floresta Formation, while the overlying Cuche Formation is much richer in fish biodiversity.
The Cuche Formation is a geological formation of the Floresta Massif, Altiplano Cundiboyacense in the Eastern Ranges of the Colombian Andes. The sequence of siltstones, shales, and sandstone beds dates to the Late Devonian and Early Carboniferous periods, and has a maximum thickness of 900 metres (3,000 ft).
