| Eomysticetidae | |
|---|---|
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Mammalia |
| Order: | Artiodactyla |
| Infraorder: | Cetacea |
| Superfamily: | Eomysticetoidea |
| Family: | † Eomysticetidae Sanders & Barnes, 2002 |
| Genera | |
Eomysticetus Contents | |
Eomysticetidae is a family of extinct mysticetes belonging to Chaeomysticeti (toothless mysticetes). It is one of two families in the basal chaeomysticete clade Eomysticetoidea (the other being Cetotheriopsidae).
Eomysticetids are united by the following combination of primitive and derived characters relative to more advanced chaeomysticetes (Balaenomorpha): zygomatic process without a supramastoid crest; reduction of the superior process of the periotic into a low ridge with anterior and posterior apices in medial or lateral view; blowholes situated ahead of the eyes; an elongated intertemporal region with long parietal and frontal exposures on the cranial vertex; elongated nasals; large coronoid processes of the mandibles; flat rostrum; laterally bowed mandibles; absence of functional teeth; and large mandibular foramina. [2]
There are seven genera of Eomysticetidae: Eomysticetus, Matapanui, Micromysticetus, Tohoraata, Tokarahia, Waharoa and Yamatocetus.
Until the early 21st century, some of the known representatives of the family were thought to belong to then-wastebasket family Cetotheriidae, including Tokarahia lophocephalus and Tohoraata waitakiensis . However, in the original description of Eomysticetus, the similarities of "Mauicetus" lophocephalus to Eomysticetus, although Sanders and Barnes (2002) stopped short of assigning "M." lophocephalus to Eomysticetidae. [3] Subsequent studies confirmed the placement of "M." lophocephalus and "M." waitakiensis in Eomysticetidae. [4] [5] [6] [7]
As members of Chaeomysticeti, eomysticetids used their baleen plates to filter krill and other planktonic organisms. Although superficially similar to thalassotherian chaeomysticetes, their large mandibular canal indicates that they were incapable of lunge-feeding as in modern-day balaenopterids. A fatty pad on the mandibular canal suggests that eomysticetids could hear underwater. [6] [7] [8] [9]
Neobalaenidae is a family of baleen whales including the extant pygmy right whale. Although traditionally considered related to balaenids, recent studies by Fordyce and Marx (2013) and Ludovic Dutoit and colleagues (2023) have recovered the living pygmy right whale as a member of Cetotheriidae, making it the only extant cetotheriid. Not all authors agree with this placement.
Janjucetus is an extinct genus of cetacean, and a basal baleen whale (Mysticeti), from the Late Oligocene around 25 million years ago (mya) off south-east Australia, containing one species J. hunderi. Unlike modern mysticetes, it possessed large teeth for gripping and shredding prey, and lacked baleen, and so was likely to have been a predator that captured large single prey animals rather than filter feeding. However, its teeth may have interlocked, much like those of the modern-day filter-feeding crabeater seal, which would have allowed some filter-feeding behaviour. Its hunting behaviour was probably similar to the modern-day leopard seal, probably eating large fish. Like baleen whales, Janjucetus could not echolocate; however, it did have unusually large eyes, and so probably had an acute sense of vision. The only specimen was found on the Jan Juc beach, where the remains of the extinct whales Mammalodon, Prosqualodon and Waipatia have also been discovered.
Mammalodon is an extinct genus of archaic baleen whale belonging to the family Mammalodontidae.
Mammalodontidae is a family of extinct whales known from the Oligocene of Australia and New Zealand.
Aetiocetus is a genus of extinct basal mysticete, or baleen whale that lived 33.9 to 23.03 million years ago, in the Oligocene in the North Pacific ocean, around Japan, Mexico, and Oregon, U.S. It was first described by Douglas Emlong in 1966 and currently contains known four species, A. cotylalveus, A. polydentatus, A. tomitai, and A. weltoni. These whales are remarkable for their retention of teeth and presence of nutrient foramina, indicating that they possessed baleen. Thus, Aetiocetus represents the transition from teeth to baleen in Oligocene mysticetes. Baleen is a highly derived character, or synapomorphy, of mysticetes, and is a keratinous structure that grows from the palate, or roof of the mouth, of the whale. The presence of baleen is inferred from the fossil record in the skull of Aetiocetus. Aetiocetus is known from both sides of the Pacific Ocean: it was first documented in Oregon, United States, but it is also known from Japan and Mexico. The genus is currently constrained to the Northern hemisphere and has little value in biostratigraphic studies of the Oligocene due to its limited occurrences across the Pacific.
Cetotheriidae is a family of baleen whales. The family is known to have existed from the Late Oligocene to the Early Pleistocene before going extinct. Although some phylogenetic studies conducted by Fordyce & Marx 2013 recovered the living pygmy right whale as a member of Cetotheriidae, making the pygmy right whale the only living cetotheriid, other authors either dispute this placement or recover Neobalaenidae as a sister group to Cetotheriidae.
The Kokoamu Greensand is a geological formation found in New Zealand. It is a fossil-bearing, late Oligocene, greensand rock unit of the eastern South Island, especially the Waitaki District of North Otago and the southern Canterbury region. The formation was named by geologist Maxwell Gage in the 1950s. In North Otago it underlies the thicker and harder Otekaike Limestone. The formation gets its green colour from the mineral glauconite which forms slowly on the ocean floor.
Eomysticetus is an extinct genus of baleen whale from the late Oligocene (Chattian) Chandler Bridge Formation of South Carolina.
Llanocetus is a genus of extinct toothed baleen whales from the Late Eocene of Antarctica. The type species, Llanocetus denticrenatus, reached gigantic proportions, with the juvenile specimen reaching an estimated 8 m (26 ft) in length; a second, unnamed species, known only from three isolated premolar teeth, reached an estimated total body length of up to 12 m (39 ft). Like other contemporary baleen whales of the Eocene, Llanocetus completely lacked baleen in its jaws. It was probably a suction feeder like the modern beaked and pygmy right whales.
Mauicetus is a genus of extinct baleen whale from the Late Oligocene of New Zealand.
Tohoraata is a genus of eomysticetid baleen whale from the Late Oligocene (Chattian) of New Zealand. There are two recognized species, T. raekohao and T. waitakiensis.
Tokarahia is a genus of eomysticetid baleen whale from the Late Oligocene (Chattian) of New Zealand. There are two recognized species, T. kauaeroa and T. lophocephalus.
Aetiocetidae is an extinct family of toothed baleen whales known from the Oligocene. The whales are from the North Pacific Ocean and ranged in size from 3 to 8 metres long. Many of the described specimens were discovered from the Upper Oligocene of the Japanese Morawan Formation, the largest known one from the Morawan's Upper tuffaceous siltstone. Other formally described extinct toothed mysticetis from this time are smaller, from 3 to 4 metres in length. Mysticeti with true baleen are seen in fossils from the Upper Oligocene. The monophyly of the family is still uncertain, as are the evolutionary relationship between the early toothed baleen whales and the early and extant edentulous baleen whales. However, the cladistic analyses of Coronodon and Mystacodon seem to indicate that Aetiocetidae and Llanocetidae are more closely related to crown Mysticeti than to Mammalodontidae, Coronodon, and Mystacodon.
Waharoa is a genus of eomysticetid baleen whale from the Late Oligocene (Chattian) of New Zealand. It was identified with the discovery of Waharoa ruwhenua by Boessenecker and Fordyce (2015), which added a new genus and species to a monophyletic family Eomysticetidae.
Matapanui is a genus of eomysticetid baleen whale from the Late Oligocene Kokoamu Greensand of New Zealand.
Horopeta is a genus of baleen whale from the Late Oligocene (Chattian) Kokoamu Greensand of New Zealand.
Coronodon is a genus of toothed (transitional) baleen whales from the Early Oligocene Ashley and Chandler Bridge formations of South Carolina. The genus contains three species: the type species C. havensteini, and additional species C. newtonorum and C. planifrons.
Whakakai is a genus of baleen whale from the Late Oligocene (Chattian) Kokoamu Greensand of New Zealand.
Sitsqwayk is a genus of baleen whale from Late Oligocene (Chattian) marine deposits in Washington state. The generic name refers to a powerful water spirit in the folklore of the Klallam that is said to bring wealth.
