| Fecampiida | |
|---|---|
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Platyhelminthes |
| Subphylum: | Rhabditophora |
| Order: | Fecampiida Rohde, Luton, Baverstock & Johnson, 1994 |
| Families | |
Fecampiida is an order of flatworms in the class Rhabditophora. It is a considerably recent clade, erected after molecular studies. [1]
The order Fecampiida, as currently defined, was erected based on molecular studies. [2] They all are parasitic organisms and are united by a similar development of the basal bodies during spermiogenesis. [3]
Three families of Fecampiida were initially classified in different flatworm orders: Urastomatidae and Genostomatidae were considered prolecithophorans, while Fecampiidae was considered a rhabdocoel. When the genus Notentera was discovered, its relationship with Fecampiidae was clear based on morphology, and both groups were united under Fecampiida. [3] Further ultrastructural studies suggested that Urastomatidae and Genostomatidae were closely related to Fecampiidae and Notenteridae, which was confirmed by molecular studies. [4]
Due to similarities in the protonephridial flame bulb, sperm and spermiogenesis, as well as the parasitic lifestyle, fecampiids were thought to be closely related to neodermatans, [3] but molecular studies revealed them to be more closely related to triclads and prolecithophorans. [4]
The flatworms, flat worms, Platyhelminthes, or platyhelminths are a phylum of relatively simple bilaterian, unsegmented, soft-bodied invertebrates. Unlike other bilaterians, they are acoelomates, and have no specialised circulatory and respiratory organs, which restricts them to having flattened shapes that allow oxygen and nutrients to pass through their bodies by diffusion. The digestive cavity has only one opening for both ingestion and egestion ; as a result, the food cannot be processed continuously.
The Turbellaria are one of the traditional sub-divisions of the phylum Platyhelminthes (flatworms), and include all the sub-groups that are not exclusively parasitic. There are about 4,500 species, which range from 1 mm (0.039 in) to large freshwater forms more than 500 mm (20 in) long or terrestrial species like Bipalium kewense which can reach 600 mm (24 in) in length. All the larger forms are flat with ribbon-like or leaf-like shapes, since their lack of respiratory and circulatory systems means that they have to rely on diffusion for internal transport of metabolites. However, many of the smaller forms are round in cross section. Most are predators, and all live in water or in moist terrestrial environments. Most forms reproduce sexually and with few exceptions all are simultaneous hermaphrodites.
Acoelomorpha is a subphylum of very simple and small soft-bodied animals with planula-like features which live in marine or brackish waters. They usually live between grains of sediment, swimming as plankton, or crawling on other organisms, such as algae and corals. With the exception of two acoel freshwater species, all known Acoelomorphs are marine.
Nannochoristidae is a family of scorpionflies with many unusual traits. It is a tiny, relict family with a single extant genus, Nannochorista, with eight species occurring in New Zealand, southeastern Australia, Tasmania, Argentina and Chile. Due to the group's distinctiveness from other scorpionflies, it is sometimes placed in its own order, the Nannomecoptera. Some studies have placed them as the closest living relatives of fleas. Most mecopteran larvae are eruciform, or shaped like caterpillars. Nannochoristid larvae, however, are elateriform, and have elongated and slender bodies. The larvae are aquatic, which is unique among mecopterans. The larvae are predatory, hunting on the beds of shallow streams, primarily on the larvae of aquatic Diptera like chironomids.
Pseudocerotidae is a family of flatworms which includes the Bedford's flatworm. Pseudocerotidae are simple organisms categorized by their oval bodies and tentacles and bright colors. They use the cilia to glide along surfaces. Most commonly referred to as marine flatworms, closely related to the orders Macrostomorpha and Lecithoepitheliata. These organisms have very complex reproductive systems, no blood systems or organs for gas exchange, a simple brain and are hermaphroditic.
Geoplanidae is a family of flatworms known commonly as land planarians or land flatworms.
Catenulida is an order of flatworms in the classical classification, or a class of flatworms in a phylogenetic approach. They are relatively small free-living flatworms, inhabiting freshwater and marine environments. There are about 100 species described worldwide, but the simple anatomy makes species distinction problematic.
Nephrozoa is a major clade of bilaterians, divided into the protostomes and the deuterostomes, containing almost all animal phyla and over a million extant species. Its sister clade is the Xenacoelomorpha. The Ambulacraria are occasionally thought to be sister to the Xenacoelomorpha, forming the Xenambulacraria as basal Deuterostomia, or basal Bilateria invalidating Nephrozoa and Deuterostomia in multiple studies. The coelom, the digestive tract and excretory organs (nephridia), and nerve cords developed in the Nephrozoa. It has been argued that, because protonephridia are only found in protostomes, they cannot be considered a synapomorphy of this group. This would make Nephrozoa an improper name, leaving Eubilateria as this clade's name.
Dimarcusidae is a family of triclads found mostly in freshwater habitats of caves, although at least one species, Rhodax evelinae, occurs in surface waters. Currently the family contains only seven species distributed in five genera, although the total number of species is thought to be much higher.
Xenacoelomorpha is a small phylum of bilaterian invertebrate animals, consisting of two sister groups: xenoturbellids and acoelomorphs. This new phylum was named in February 2011 and suggested based on morphological synapomorphies, which was then confirmed by phylogenomic analyses of molecular data.
Cavernicola is a suborder of planarians found mostly in freshwater habitats of caves, although some species occur on the surface.
Geoplaninae is a subfamily of land planarians endemic to the Neotropical region. Members of this family are sometimes referred to as the Neotropical land planarians. However, one species, Obama nungara has been introduced in Europe.
Obama is a genus of land planarians from South America. It contains several species adapted to human-disturbed environments, including the only invasive land planarian native to the Neotropical realm, Obama nungara, which has been accidentally introduced in Europe.
Neodermata is a clade of rhabditophoran flatworms containing the parasitic groups Trematoda, Monogenea and Cestoda.
Rhabditophora is a class of flatworms. It includes all parasitic flatworms and most free-living species that were previously grouped in the now obsolete class Turbellaria. Therefore, it contains the majority of the species in the phylum Platyhelminthes, excluding only the catenulids, to which they appear to be the sister group.
Rhabdocoela is an order of flatworms in the class Rhabditophora with about 1700 species described worldwide. The order was first described in 1831 by Christian Gottfried Ehrenberg. Most of rhabdocoels are free-living organisms, but some live symbiotically with other animals.
Proseriata is an order of free-living flatworms in the class Rhabditophora with over 400 species described worldwide.
Trepaxonemata is a subclass of the Platyhelminthes or flatworms. It includes all parasitic flatworms and several free-living species that were previously grouped in the now obsolete class Turbellaria. Therefore, it contains the majority of species in the phylum Platyhelminthes, excluding the Catenulida, and the Macrostomorpha.
Kalyptorhynchia is a suborder of rhabdocoel flatworms. It contains almost 600 species and has a cosmopolitan distribution.
Dalytyphloplanida is a suborder of rhabdocoel flatworms. It contains about 1000 species and has a cosmopolitan distribution in both marine and freshwater environments, with several groups having commensal or parasitic lifestyles.
