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| Geomyoidea Temporal range: Early Eocene - Recent | |
|---|---|
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| Kangaroo rat | |
Scientific classification  | |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Mammalia |
| Order: | Rodentia |
| Suborder: | Castorimorpha |
| Superfamily: | Geomyoidea Bonaparte, 1845 |
| Families | |
†Eomyidae Contents | |
Geomyoidea is a superfamily of rodent that contains the pocket gophers (Geomyidae), the kangaroo rats and mice (Heteromyidae), and their fossil relatives.
Although dissimilar in overall appearance, gophers have been united with kangaroo rats into a common superfamily for a considerable time. The superfamily Geomyoidea is among the few superfamilial relationships in rodents that is not subject to much controversy. Overall morphology, the fossil record, molecular analyses, and biogeography all support this relationship.
Geomyoids are most noticeably characterized by the position of the infraorbital canal. Unlike all other rodents who have the opening of the infraorbital canal facing forward, geomyoids have an infraorbital canal that faces to the side. Instead of passing through the zygoma, the infraorbital canal of geomyoids has moved to the side of the snout. This condition is so pronounced and the snout so narrow in heteromyids that the infraorbital canals from either side connect. Essentially, if the skull of a heteromyid is viewed from the side, the viewer can see directly through it.
Modern geomyoids are mostly restricted to North America, but some representatives have extended their range into South America since the Great American Interchange. Fossil taxa are known from throughout Laurasia.
Geomyoids have been considered to be either sciuromorphous or myomorphous depending on the authority. The masseter muscle does not pass through the infraorbital canal; it cannot due to the position of the canal. Some authorities consider the geomyoids related to squirrels, beavers, and mountain beavers on this basis.
The masseter muscle does attach directly behind the zygomatic arch in a manner very different from sciuromorphs. Some authorities consider geomyoids myomorphs based on this feature. This suggests they may be related to mice, jerboas, and perhaps dormice.
The family †Eomyidae is alternatively referred to as a member of the superfamily Geomyoidea or as a separate superfamily (†Eomyoidea) within the shared infraorder "Geomorpha" (which is also used for a genus of shield-bugs). †Florentiamyidae and †Heliscomyidae are usually placed within the superfamily Geomyoidea regardless of if eomyids are treated as a separate superfamily or not (Korth et al., 1991). McKenna and Bell (1997) do not recognize heliscomyids as a distinct family, placing the one or two heliscomyid genera in Geomyoidea incertae sedis. Sometimes the pocket gophers and heteromyids are placed as separate subfamilies within a single family (Geomyidae). These subfamilies are Geomyinae and Heteromyinae respectively.
Cladogram showing interrelationships among geomyoid families following Korth et al. (1991):
| Geomyoidea |
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Kangaroo rats, small mostly nocturnal rodents of genus Dipodomys, are native to arid areas of western North America. The common name derives from their bipedal form. They hop in a manner similar to the much larger kangaroo, but developed this mode of locomotion independently, like several other clades of rodents.
Heteromyidae is a family of rodents consisting of kangaroo rats, kangaroo mice, pocket mice and spiny pocket mice. Most heteromyids live in complex burrows within the deserts and grasslands of western North America, though species within the genus Heteromys are also found in forests and their range extends down as far as northern South America. They feed mostly on seeds and other plant parts, which they carry in their fur-lined cheek pouches to their burrows.
Mammalia is a class of animal within the phylum Chordata. Mammal classification has been through several iterations since Carl Linnaeus initially defined the class. No classification system is universally accepted; McKenna & Bell (1997) and Wilson & Reader (2005) provide useful recent compendiums. Many earlier ideas from Linnaeus et al. have been completely abandoned by modern taxonomists, among these are the idea that bats are related to birds or that humans represent a group outside of other living things. Competing ideas about the relationships of mammal orders do persist and are currently in development. Most significantly in recent years, cladistic thinking has led to an effort to ensure that all taxonomic designations represent monophyletic groups. The field has also seen a recent surge in interest and modification due to the results of molecular phylogenetics.
Caviomorpha is the rodent infraorder or parvorder that unites all New World hystricognaths. It is supported by both fossil and molecular evidence. The Caviomorpha was for a time considered to be a separate order outside the Rodentia, but is now accepted as a genuine part of the rodents. Caviomorphs include the extinct Heptaxodontidae, the extinct Josephoartigasia monesi and extant families of chinchilla rats, hutias, guinea pigs and the capybara, chinchillas and viscachas, tuco-tucos, agoutis, pacas, pacaranas, spiny rats, New World porcupines, coypu and octodonts.
The Hystricognathi are an infraorder of rodents, distinguished from other rodents by the bone structure of their skulls. The masseter medialis passes partially through a hole below each eye socket and connects to the bone on the opposite side. This, together with their lack of an infraorbital plate and the relative size of the infraorbital foramen, distinguishes hystricognaths from other rodent groups.
Sciuromorpha ("squirrel-like") is a rodent clade that includes several different rodent families. It includes all members of the Sciuridae as well as the mountain beaver species.
The suborder Myomorpha contains 1,524 species of mouse-like rodents, nearly a quarter of all mammal species. Included are mice, rats, gerbils, hamsters, lemmings, and voles. They are grouped according to the structure of their jaws and molar teeth. They are characterized by their myomorphous zygomasseteric system, which means that both their medial and lateral masseter muscles are displaced forward, making them adept at gnawing. As in the hystricognathous rodents, the medial masseter muscle goes through the eye socket, a feature unique among mammals. Myomorphs are found worldwide in almost all land habitats. They are usually nocturnal seed-eaters.
The blesmols, also known as mole-rats, or African mole-rats, are burrowing rodents of the family Bathyergidae. They represent a distinct evolution of a subterranean life among rodents much like the pocket gophers of North America, the tuco-tucos in South America, or the Spalacidae from Eurasia.
Castorimorpha is the suborder of rodents containing the beavers and the kangaroo rats. A 2017 study using retroposon markers indicated that they are most closely related to the Anomaluromorpha and Myomorpha.
Perognathinae is a subfamily of rodents consisting of two genera of pocket mice. Most species live in complex burrows within the deserts and grasslands of western North America, They feed mostly on seeds and other plant parts, which they carry in their fur-lined cheek pouches to their burrows.
The zygomasseteric system in rodents is the anatomical arrangement of the masseter muscle of the jaw and the zygomatic arch of the skull. The anteroposterior or propalinal (front-to-back) motion of the rodent jaw is enabled by an extension of the zygomatic arch and the division of the masseter into a superficial, lateral and medial muscle. The four main types are described as protrogomorphous, sciuromorphous, hystricomorphous, and myomorphous.
Eomyidae is a family of extinct rodents from North America and Eurasia related to modern day pocket gophers and kangaroo rats. They are known from the Middle Eocene to the Late Miocene in North America and from the Late Eocene to the Pleistocene in Eurasia. Eomyids were generally small, but occasionally large, and tended to be squirrel-like in form and habits. The family includes the earliest known gliding rodent, Eomys quercyi.
Heliscomyidae is a family of extinct rodents from the mid-Tertiary of North America related to pocket gophers and kangaroo rats and their relatives. The family contains four genera, Apletotomeus, Heliscomys, Passaliscomys, and Tylionomys. McKenna and Bell (1997) placed the first two genera in synonymy, with Heliscomys the senior synonym.
Merriam's kangaroo rat is a species of rodent in the family Heteromyidae. The species name commemorates Clinton Hart Merriam. It is found in the Upper and Lower Sonoran life zones of the southwestern United States, Baja California, and northern Mexico.
The narrow-faced kangaroo rat is a species of rodent in the family Heteromyidae. It is endemic to California in the United States.
Rodents are mammals of the order Rodentia, which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are native to all major land masses except for New Zealand, Antarctica, and several oceanic islands, though they have subsequently been introduced to most of these land masses by human activity.
Handbook of the Mammals of the World (HMW) is a book series from the publisher Lynx Edicions. The nine volumes were published from 2009 to 2019. Each mammal family is assessed in a full text introduction with photographs and each species has a text account with a distribution map and illustrations on a plate. This is the second major project by Lynx Edicions since the release of the Handbook of the Birds of the World in 1992. The chief editors are Russell Mittermeier and Don E. Wilson in association with Conservation International, the Texas A&M University and the IUCN. Don E. Wilson is also editor of the reference work Mammal Species of the World.
Cheek pouches are pockets on both sides of the head of some mammals between the jaw and the cheek. They can be found on mammals including the platypus, some rodents, and most monkeys, as well as the marsupial koala. The cheek pouches of chipmunks can reach the size of their body when full.