Heliscomyidae Temporal range: | |
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Fossil | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Rodentia |
Superfamily: | Geomyoidea |
Family: | † Heliscomyidae Korth, Wahlert, and Emry, 1991 |
Genera | |
Heliscomyidae is a family of extinct rodents from the mid-Tertiary of North America related to pocket gophers (family Geomyidae) and kangaroo rats and their relatives (family Heteromyidae). The family contains four genera, Apletotomeus , Heliscomys , Passaliscomys , and Tylionomys (Korth et al., 1991; Korth and Eaton, 2004; Korth and Branciforte, 2007). McKenna and Bell (1997) placed the first two genera in synonymy, with Heliscomys as the senior synonym.
Heliscomyids are distinguished from other geomyoid rodents by several characteristics of the skull including the fusion of three cranial foramina, elongation of the incisive foramina, and an unusual position of the mental foramen (Korth et al., 1991).
Heliscomyidae is a member of the clade Geomyoidea, a group of rodents that also includes the families †Eomyidae, †Florentiamyidae, Heteromyidae, and Geomyidae. The following cladogram showing interrelationships among geomyoid families follows Korth et al. (1991):
Geomyoidea |
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Castoridae is a family of rodents that cointains the two living species of beavers and their fossil relatives. A formerly diverse group, only a single genus is extant today, Castor. Two other genera of "giant beavers", Castoroides and Trogontherium, became extinct in the Late Pleistocene.
The family Aplodontiidae also known as Aplodontidae, Haplodontiidae or Haploodontini is traditionally classified as the sole extant family of the suborder Protrogomorpha. It may be the sister family of the Sciuridae. There are fossils from the Oligocene until Miocene in Asia, from Oligocene in Europe and from the Oligocene until the present in North America, where there is the only living species: the mountain beaver.
Simplicidentata is a group of mammals that includes the rodents and their closest extinct relatives. The term has historically been used as an alternative to Rodentia, contrasting the rodents with their close relatives the lagomorphs. However, Simplicidentata is now defined as including all members of Glires that share a more recent common ancestor with living rodents than with living lagomorphs. Thus, Simplicidentata is a total group that is more inclusive than Rodentia, a crown group that includes all living rodents, their last common ancestor, and all its descendants. Under this definition, the loss of the second pair of upper incisors is a synapomorphic feature of Simplicidentata. The loss of the second upper premolar (P2) has also been considered as synapomorphic for Simplicidentata, but the primitive simplicidentate Sinomylus does have a P2.
Castorimorpha is the suborder of rodents containing the beavers and the kangaroo rats. A 2017 study using retroposon markers indicated that they are most closely related to the Anomaluromorpha and Myomorpha.
Geomyoidea is a superfamily of rodent that contains the pocket gophers (Geomyidae), the kangaroo rats and mice (Heteromyidae), and their fossil relatives.
Eomyidae is a family of extinct rodents from North America and Eurasia related to modern day pocket gophers and kangaroo rats. They are known from the Middle Eocene to the Late Miocene in North America and from the Late Eocene to the Pleistocene in Eurasia. Eomyids were generally small, but occasionally large, and tended to be squirrel-like in form and habits. The family includes the earliest known gliding rodent, Eomys quercyi.
Mojavemyinae is an extinct subfamily of geomyoid rodents. When Mojavemyinae was named, it was not assigned to a family, with a relationship to either Geomyidae or Heteromyidae considered equally likely, but consensus places the type genus, Mojavemys, in Geomyidae.
Eurymylidae is a family of extinct simplicidentates. Most authorities consider them to be basal to all modern rodents and may have been the ancestral stock whence the most recent common ancestor of all modern rodents arose. However, the more completely known eurymylids, including Eurymylus, Heomys, Matutinia, and Rhombomylus, appear to represent a monophyletic side branch not directly ancestral to rodents. Huang et al. (2004) have argued that Hanomys, Matutinia, and Rhombomylus form a clade characterized by distinctive features of the skull and dentition that should be recognized as a separate family, Rhombomylidae. Eurymylids are only known from Asia.
Armintomys is an extinct genus of rodent from North America related to jerboas and jumping mice. It is the only genus in the family Armintomyidae. It lived during the early Eocene, and is the oldest known example of a hystricomorphous zygomasseteric dentition. In addition, Armintomys is also the oldest known rodent that had an incisor enamel transition from pauciserial to uniserial. Its remains have only been found in the Wind River Basin in Wyoming, and could be found there during the species' existence on Earth. It was previously assumed that Armintomys belonged to the Dipodoidea family, but has since been understood to have been part of an early radiation of dipodoid rodents, but was not directly ancestral to any later dipodoids, thus it was recategorized into its own family.
The Tsaganomyidae are an extinct family of rodents from Asia. It contains three genera. Tsaganomyids are generally considered to be related to the Hystricognathi. Members of Tsaganomyidae were fossorial (digging) rodents that probably used their incisor teeth to dig like some living mole rats.
Sciurini is a tribe that includes about forty species of squirrels, mostly from the Americas. It includes five living genera—the American dwarf squirrels, Microsciurus; the Bornean Rheithrosciurus; the widespread American and Eurasian tree squirrels of the genus Sciurus, which includes some of the best known squirrel species; the Central American Syntheosciurus; and the American pine squirrels, Tamiasciurus. Like other arboreal squirrels, they are sometimes referred to as tree squirrels.
The Mylagaulidae or mylagaulids are an extinct clade of sciuromorph rodents nested within the family Aplodontiidae. They are known from the Neogene of North America and China. The oldest member is the Late Oligocene Trilaccogaulus montanensis that lived some 29 million years ago (Mya), and the youngest was Ceratogaulus hatcheri—formerly in the invalid genus "Epigaulus" —which was found barely into the Pliocene, some 5 Mya.
Cephalomyidae is an extinct family of caviomorph rodents from South America. The specific relationships of the family are uncertain, and affinities to both chinchilloid and cavioid rodents have been supported. Most recently, Kramarz in 2005 performed a phylogenetic analysis supporting a relationship to the Cavioidea, as represented by Eocardiidae, although more recent analyses have placed them among the chinchilloids as relatives of the giant neoepiblemid rodents. McKenna and Bell (1997) questioned the validity of the family, placing the cephalomyid genera then known in Dasyproctidae, but Kramarz (2001) subsequently reasserted the distinctiveness of cephalomyids.
Mesotheriidae is an extinct family of notoungulate mammals known from the Oligocene through the Pleistocene of South America. Mesotheriids were small to medium-sized herbivorous mammals adapted for digging.
?Oryzomys pliocaenicus is a fossil rodent from the Hemphillian of Kansas, central United States. It is known from a single mandible with the back part missing. All three molars are present, but very worn. Together, the molars are 3.6 mm long. The fossil was discovered in 1935 and described in 1939 as a possible species of Oryzomys. Later authors doubted this allocation and suggested that it may instead belong in Bensonomys or Jacobsomys, but the material may not allow a definite identification.
Paronychomys is an extinct genus of Cricetidae that existed in Arizona during the Hemphillian period.
Brontotheriidae is a family of extinct mammals belonging to the order Perissodactyla, the order that includes horses, rhinoceroses, and tapirs. Superficially, they looked rather like rhinos, although they were actually more closely related to horses; Equidae and Brontotheriidae make up the suborder Hippomorpha. They lived around 56–34 million years ago, until the very close of the Eocene.
Apeomyoides savagei is a fossil rodent from the Miocene of the United States, the only species in the genus Apeomyoides. It is known from fragmentary jaws and isolated teeth from a site in the early Barstovian, around 15–16 million years ago, of Nevada. Together with other species from scattered localities in the United States, Japan, and Europe, Apeomyoides is classified in the subfamily Apeomyinae of the extinct rodent family Eomyidae. Apeomyines are a rare but widespread group that may have been adapted to a relatively dry habitat.
The Washakie Formation is a geologic formation in northern Colorado and southern Wyoming. It preserves many mammal, bird, reptile and other fossils dating back to the Lutetian stage of the Eocene within the Paleogene period. The sediments fall in the Bridgerian and Uintan stages of the NALMA classification.
This paleomammalogy list records new fossil mammal taxa that were described during the year 2009, as well as notes other significant paleomammalogy discoveries and events which occurred during that year.