Gigantoproductus giganteus

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Gigantoproductus giganteus
Temporal range: Carboniferous
~339.4–318.1  Ma
Gigantoproductus giganteus Naturalis.JPG
Scientific classification
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G. giganteus
Binomial name
Gigantoproductus giganteus
(Sowerby, 1822) [1]

Gigantoproductus giganteus ("Gigantic giant Productus " [2] ) is an extinct species of brachiopods in the family Monticuliferidae, [3] known only from its fossil remains. It was a marine invertebrate found on the seabed in shallow seas. It evolved during the Carboniferous period and it is believed to be the largest brachiopod that has ever existed. [2] [1]

Contents

Description

Gigantoproductus giganteus was a large brachiopod that superficially resembled a cockle. Fossils of this species have been found with shell widths of 30 centimetres (12 in). [2] and more than 35 cm (14 in). [4] It had a pair of thick dome-shaped valves joined by a hinge. The valves had a small number of broad ribs that radiated from a thick umbo and there were large wing-shaped ears of calcareous material on either side. The valves were held together by a central strong adductor muscle which left a scar on the inside of the valves. The ventral valve, also known as the pedicle valve, was covered with spines on the outside. The inside of this valve was rough, being covered by numerous cone-shaped protrusions. [5] These are visible in an internal mould of the brachiopod, a cast fossil which has been formed when a hole in sediment left by the soft tissues of the dead organism was later infiltrated by mineral matter. [6]

Distribution and habitat

The fossil record suggests that Gigantoproductus giganteus was common between about 345 and 328 million years ago. Fossils have been found in Europe, Asia and North Africa. [1] The morphology of the shell makes it likely that it lived on the seabed, partially buried in sand or mud, in shallow water in locations with strong waves and currents. Here its large size, its heavy weight, its ears and spines would have helped provide stability, preventing it from rolling around. [7]

Biology

These brachiopods were photoautotrophic. [2] Like other brachiopods, Gigantoproductus giganteus was a filter feeder, using its lophophore, a specialist feeding organ, to extract planktonic particles from the water. Reproduction was likely to have involved release of gametes into the water column. [1]

Related Research Articles

<span class="mw-page-title-main">Carboniferous</span> Fifth period of the Paleozoic Era, 359–299 million years ago

The Carboniferous is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period 358.9 million years ago (Mya), to the beginning of the Permian Period, 298.9 million years ago. The name Carboniferous means "coal-bearing", from the Latin carbō ("coal") and ferō, and refers to the many coal beds formed globally during that time.

<span class="mw-page-title-main">Bivalvia</span> Class of molluscs

Bivalvia, in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, bivalves have no head and they lack some usual molluscan organs, like the radula and the odontophore. The class includes the clams, oysters, cockles, mussels, scallops, and numerous other families that live in saltwater, as well as a number of families that live in freshwater. The majority are filter feeders. The gills have evolved into ctenidia, specialised organs for feeding and breathing. Most bivalves bury themselves in sediment, where they are relatively safe from predation. Others lie on the sea floor or attach themselves to rocks or other hard surfaces. Some bivalves, such as the scallops and file shells, can swim. The shipworms bore into wood, clay, or stone and live inside these substances.

<span class="mw-page-title-main">Bear Gulch Limestone</span>

The Bear Gulch Limestone is a limestone-rich geological lens in central Montana, renowned for the quality of its late Mississippian-aged fossils. It is exposed over a number of outcrops northeast of the Big Snowy Mountains, and is often considered a component of the more widespread Heath Formation. The Bear Gulch Limestone reconstructs a diverse, though isolated, marine ecosystem which developed near the end of the Serpukhovian age. It is a lagerstätte, a particular type of rock unit with exceptional fossil preservation of both articulated skeletons and soft tissues. Bear Gulch fossils include a variety of fish, invertebrates, and algae occupying a number of different habitats within a preserved shallow bay.

<i>Lingula</i> (brachiopod) Genus of brachiopods within the class Lingulata

Lingula is a genus of brachiopods within the class Lingulata. Lingula or forms very close in appearance have existed possibly since the Cambrian. Like its relatives, it has two unadorned organo-phosphatic valves and a long fleshy stalk. Lingula lives in burrows in barren sandy coastal seafloor and feeds by filtering detritus from the water. It can be detected by a short row of three openings through which it takes in water (sides) and expels it again (middle).

<span class="mw-page-title-main">Craniidae</span> Family of shelled animals

The Craniidae are a family of brachiopods, the only surviving members of the subphylum Craniiformea. They are the only members of the order Craniida, the monotypic suborder Craniidina, and the superfamily Cranioidea; consequently, the latter two taxa are at present redundant and rarely used.There are three living genera within Craniidae: Neoancistrocrania, Novocrania, and Valdiviathyris. As adults, craniids either live freely on the ocean floor or, more commonly, cement themselves onto a hard object with all or part of the ventral valve.

<i>Terebratula</i>

Terebratula is a modern genus of brachiopods with a fossil record dating back to the Late Devonian. These brachiopods are stationary epifaunal suspension feeders and have a worldwide distribution.

<i>Atrypa</i> Genus of brachiopod

Atrypa is a genus of brachiopod with shells round to short egg-shaped, covered with many fine radial ridges, that split further out and growth lines perpendicular to the costae and 2-3 times wider spaced. The pedunculate valve is a little convex, but tends to level out or even become slightly concave toward the anterior margin. The brachial valve is highly convex. There is no interarea in either valve. Atrypa was a cosmopolitan and occurred from the late Lower Silurian (Telychian) to the early Upper Devonian (Frasnian). Other sources expand the range from the Late Ordovician to Carboniferous, approximately from 449 to 336 Ma. A proposed new species, A. harrisi, was found in the trilobite-rich Floresta Formation in Boyacá, Colombia.

<span class="mw-page-title-main">Strophomenida</span> Extinct order of brachiopods

Strophomenida is an extinct order of articulate brachiopods which lived from the lower Ordovician period to the mid Carboniferous period. Strophomenida is part of the extinct class Strophomenata, and was the largest known order of brachiopods, encompassing over 400 genera. Some of the largest and heaviest known brachiopod species belong to this class. Strophomenids were among the most diverse and abundant brachiopods during the Ordovician, but their diversity was strongly impacted at the Late Ordovician mass extinction. Survivors rediversified into new morphologies in the Silurian, only to be impacted once again at the Late Devonian mass extinction. However, they still survived till the mid Carboniferous.

<span class="mw-page-title-main">Brachiopod</span> Phylum of marine animals also known as lamp shells

Brachiopods, phylum Brachiopoda, are a phylum of trochozoan animals that have hard "valves" (shells) on the upper and lower surfaces, unlike the left and right arrangement in bivalve molluscs. Brachiopod valves are hinged at the rear end, while the front can be opened for feeding or closed for protection. Two major categories are traditionally recognized, articulate and inarticulate brachiopods. The word "articulate" is used to describe the tooth-and-groove structures of the valve-hinge which is present in the articulate group, and absent from the inarticulate group. This is the leading diagnostic skeletal feature, by which the two main groups can be readily distinguished as fossils. Articulate brachiopods have toothed hinges and simple, vertically-oriented opening and closing muscles. Conversely, inarticulate brachiopods have weak, untoothed hinges and a more complex system of vertical and oblique (diagonal) muscles used to keep the two valves aligned. In many brachiopods, a stalk-like pedicle projects from an opening near the hinge of one of the valves, known as the pedicle or ventral valve. The pedicle, when present, keeps the animal anchored to the seabed but clear of sediment which would obstruct the opening.

<span class="mw-page-title-main">Evolution of brachiopods</span> The origin and diversification of brachiopods through geologic time

The origin of the brachiopods is uncertain; they either arose from reduction of a multi-plated tubular organism, or from the folding of a slug-like organism with a protective shell on either end. Since their Cambrian origin, the phylum rose to a Palaeozoic dominance, but dwindled during the Mesozoic.

<i>Gigantoproductus</i>

Gigantoproductus is a genus of extinct brachiopods in the order Productida and the family Monticuliferidae. The species were the largest of the carboniferous brachiopods, with the largest known species reaching 30 cm (12 in) in shell width. Such huge invertebrates appeared in the Mississippian as the proportion of oxygen in the atmosphere began to rise. The earliest members of the Productida date back to the Silurian period, and Gigantoproductus is known to have existed between 339.4 to 318.1 million years ago, during the Carboniferous period. As fossils, their shells occur within a limestone matrix.

The cephalopods have a long geological history, with the first nautiloids found in late Cambrian strata, and purported stem-group representatives present in the earliest Cambrian lagerstätten.

<span class="mw-page-title-main">Rhynchonelliformea</span> Subphylum of brachiopods

Rhynchonelliformea is a major subphylum and clade of brachiopods. It is roughly equivalent to the former class Articulata, which was used previously in brachiopod taxonomy up until the 1990s. These so-called articulated brachiopods have many anatomical differences relative to "inarticulate" brachiopods of the subphyla Linguliformea and Craniformea. Articulates have hard calcium carbonate shells with tongue-and-groove hinge articulations and separate sets of simple opening and closing muscles.

<i>Whatcheeria</i> Extinct genus of tetrapods

Whatcheeria is an extinct genus of early tetrapod from the Mississippian of Iowa. Fossils have been found in 340 million year old fissure fill deposits in the town of Delta. The type species, Whatcheeria deltae was named in 1995. It is classified within the family Whatcheeriidae, along with the closely related Pederpes and possibly Ossinodus.

<span class="mw-page-title-main">Paleontology in New Mexico</span>

Paleontology in New Mexico refers to paleontological research occurring within or conducted by people from the U.S. state of New Mexico. The fossil record of New Mexico is exceptionally complete and spans almost the entire stratigraphic column. More than 3,300 different kinds of fossil organisms have been found in the state. Of these more than 700 of these were new to science and more than 100 of those were type species for new genera. During the early Paleozoic, southern and western New Mexico were submerged by a warm shallow sea that would come to be home to creatures including brachiopods, bryozoans, cartilaginous fishes, corals, graptolites, nautiloids, placoderms, and trilobites. During the Ordovician the state was home to algal reefs up to 300 feet high. During the Carboniferous, a richly vegetated island chain emerged from the local sea. Coral reefs formed in the state's seas while terrestrial regions of the state dried and were home to sand dunes. Local wildlife included Edaphosaurus, Ophiacodon, and Sphenacodon.

<i>Argyrotheca</i>

Argyrotheca is a genus of very small to minute lampshells. All species share a large pedicel opening, one ridge on the inside of the pedunculate valve, pits in a diamond pattern on the inside of both valves, and without radial ridges that end in tubercles. It occurs in depths between 6 and 1300 m. It is known since the latest Cretaceous.

<span class="mw-page-title-main">Siphonotretida</span> Extinct order of marine lamp shells

Siphonotretida is an extinct order of linguliform brachiopods in the class Lingulata. The order is equivalent to the sole superfamily Siphonotretoidea, itself containing the sole family Siphonotretidae. Siphonotretoids were originally named as a superfamily of Acrotretida, before being raised to their own order.

<span class="mw-page-title-main">Kutorginata</span> Extinct genus of shelled animals

Kutorginates (Kutorginata) are an extinct class of early rhynchonelliform ("articulate") brachiopods. The class contains only a single order, Kutorginida (kutorginides). Kutorginides were among the earliest rhynchonelliforms, restricted to the lower-middle part of the Cambrian Period.

<span class="mw-page-title-main">Orthotetida</span> Extinct order of marine lamp shells

The orthotetides (Orthotetida) are an extinct order of brachiopods in the class Strophomenata. Though not particularly diverse or abundant relative to strophomenides (Strophomenida) or productides (Productida), orthotetides were nevertheless the longest-lasting order of strophomenates, surviving from the Middle Ordovician (“Llanvirn”) up until the Late Permian. Externally, many orthotetides are difficult to distinguish from strophomenides. Most fundamental differences between the two orders are internal: orthotetides have more elaborate cardinal processes and a greater diversity of shell microstructure.

Pugnoides is an extinct genus of brachiopod belonging to the order Rhynchonellida and family Petasmariidae. Specimens have been found in Devonian to Permian beds in North America, Asia, Europe, western Australia, New Zealand,and New Zealand. The genus was particularly widespread in the Visean.

References

  1. 1 2 3 4 Lee Davies. "Gigantoproductus giganteus". Natural History Museum. Retrieved 2013-02-21.
  2. 1 2 3 4 George R. McGhee, Jr. (2019). Convergent Evolution on Earth. Lessons for the Search for Extraterrestrial Life. MIT Press. p. 47. ISBN   9780262354189 . Retrieved 2022-08-23.
  3. A. Williams, S. J. Carlson, C. H. C. Brunton, L. E. Holmer, L. E. Popov, M. Mergl, J. R. Laurie, M. G. Bassett, L. R. M. Cocks, J.-Y. Rong, S. S. Lazarev, R. E. Grant, P. R. Racheboeuf, Y.-G. Jin, B. R. Wardlaw, D. A. T. Harper, and A. D. Wright and M. 2000. Part H Brachiopoda (revised): Volumes 2 & 3, Linguliformea, Craniiformea, Rhynchonelliformea (part). Treatise on Invertebrate Paleontology 1-919 [W. Kiessling/W. Kiessling/W. Kiessling]
  4. Vojtěch Turek, J. Marek, Josef Beneš (1989). Fossils of the World. A Comprehensive Practical Guide to Collecting and Studying Fossils. Arch Cape Press. p. 146. ISBN   9780517679043 . Retrieved 2022-08-23.{{cite book}}: CS1 maint: multiple names: authors list (link)
  5. Tom Yancey. "Gigantoproductus giganteus (Sowerby, 1822)". Carboniferous fossils of Russia. Retrieved 2013-02-21.
  6. Megan Beckett. "Fossils: Fossil types". Siyavula: Life Sciences Grade 10. Retrieved 2013-02-23.
  7. Ferguson, J. (1978). "Some aspects of the ecology and growth of the carboniferous gigantoproductids". Proceedings of the Yorkshire Geological Society. 42: 41–54. doi:10.1144/pygs.42.1.41.