Ginkgo dissecta Temporal range: | |
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G. dissecta leaf | |
Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
(unranked): | Gymnosperms |
Division: | Ginkgophyta |
Class: | Ginkgoopsida |
Order: | Ginkgoales |
Family: | Ginkgoaceae |
Genus: | Ginkgo |
Species: | †G. dissecta |
Binomial name | |
†Ginkgo dissecta Mustoe, 2002 | |
Ginkgo dissecta is an extinct ginkgo species in the family Ginkgoaceae described from a series of isolated fossil leaves. [1] The species is known from Early Eocene [2] sediments exposed in the province of British Columbia, Canada, and Washington, USA. It is one of two Ginkgo species found at the Washington and British Columbia sites. [1]
Ginkgo dissecta is represented by a group of fossil specimens from four different geologic formations. The type locality for the species is at the Ypresian [2] McAbee Fossil Beds, near Cache Creek, British Columbia, in the Tranquille Formation belonging to the Kamloops Group. G. dissecta is also known from the similarly aged sites of the Klondike Mountain Formation, which crop out around the town of Republic, Ferry County, Washington. [1] At least one specimen has been recovered from the fossil sites around the town of Princeton, British Columbia, [1] and a fourth occurrence for the species was reported from the "Falkland site" near the town of Falkland, British Columbia, also part of the Tranquille Formation. [3]
The type specimens for G. dissecta include two leaf fossils, a holotype and a paratype, both from the McAbee Fossil Beds. The holotype leaf is numbered number WWU-GK-020 and the paratype is number WWU-GK-008, both of which are currently preserved in the paleobotanical collections of Western Washington University in Bellingham, Washington. [1] In addition to the two type specimens, the Western Washington University collections have 39 other fossils from the McAbee site and one from the Princeton fossil sites. For the species description one fossil, SR 96-09-01, was borrowed from the Stonerose Interpretive Center in Republic, Washington. The specimens were studied by paleobotanist George Mustoe of the Western Washington University Geology Department. Mustoe published his 2002 type description for G. dissecta in the Canadian Journal of Botany . The etymology of the chosen specific name dissecta was not identified by Mustoe in the type description, but he noted it is a formalization of the name which had been first used in 1974 in an unpublished thesis by Verschoor. [1]
Leaf widths of G. dissecta range between 50 and 80 mm (2.0 and 3.1 in) and petioles reaching up to 40 mm (1.6 in) in length. In contrast to the two-lobed structure of modern Ginkgo biloba leaves, the morphology of G. dissecta is four-lobed. These lobes are separated by deep notches which almost reach the leaf base, while additional smaller notches may be present in the upper portions of the lobes. The vein structure of G. dissecta is composed of numerous forking veins which start at the base of the leaf. This forking is different from the vein structure of G. biloba in which all the veins fork from a single vein along the edge of the blade. [1] The overall vein density in G. dissecta is 12–14 veins per centimeter, which is lower than the reported average of 16–17 veins per centimeter in G. biloba. G. dissecta is superficially similar to the older species G. digitata and G. huttonii known from Jurassic fossils. However, both G. digitata and G. huttoni have leaves which are typically divided into six lobes rather than four and both of them have distinct trichomes, hairs, on the underside of the leaves. [1]
Ginkgoales are a gymnosperm order containing only one extant species: Ginkgo biloba, the ginkgo tree. It is monotypic, within the class Ginkgoopsida, which itself is monotypic within the division Ginkgophyta. The order includes five families, of which only Ginkgoaceae remains extant.
Florissantia is an extinct genus of flowering plants in the Malvaceae subfamily Sterculioideae known from western North America and far eastern Asia. Flower, fruit, and pollen compression fossils have been found in formations ranging between the Early Eocene through to the Early Oligocene periods. The type species is Florissantia speirii and three additional species are known, Florissantia ashwillii, Florissantia quilchenensis, and Florissantia sikhote-alinensis.
Tilia johnsoni is an extinct species of flowering plant in the family Malvaceae that, as a member of the genus Tilia, is related to modern lindens. The species is known from fossil leaves found in the early Eocene deposits of northern Washington state, United States and a similar aged formation in British Columbia, Canada.
Dillhoffia is an extinct monotypic genus of flowering plant with a single species, Dillhoffia cachensis known from Ypresian age Eocene fossils found in British Columbia, Canada, and Washington, USA. The genus and species were described from fifteen specimens found in an unnamed formation belonging to the Kamloops group shales; and two specimens from the Klondike Mountain Formation. The unnamed formation outcrops at the McAbee Fossil Beds near Cache Creek, BC, which is designated the type locality while the two U.S. specimens were recovered from the Tom Thumb Tuff member of the Klondike Mountain Formation in Republic, Washington. It is of interest to note that of the Okanagan highlands fossil sites Dillhoffia is only known from two locations, and is absent or has not been identified from the others.
Acer stewarti is an extinct maple species in the family Sapindaceae described from a series of fossil leaves and samaras. The species is solely known from the Early Eocene sediments exposed in south central British Columbia, Canada adjacent to northeast Washington state, United States. It is one of only two species belonging to the extinct section Stewarta.
Acer rousei is an extinct maple species in the family Sapindaceae described from a series of isolated fossil samaras. The species is solely known from the Early Eocene sediments exposed in south central British Columbia, Canada adjacent to northeast Washington state in the United States. It is the type species for the extinct monotypic section Rousea.
Acer browni is an extinct maple species in the family Sapindaceae described from a series of isolated fossil leaves and samaras. The species is known from the early to middle Miocene sediments exposed in Western Oregon, Washington state, USA and Northern Graham Island, Haida Gwaii, Canada. It is one of several extinct species placed in the living section Parviflora.
Fothergilla malloryi is an extinct species of flowering plant in the family Hamamelidaceae known from fossil leaves found in the early Eocene Klondike Mountain Formation deposits of northern Washington state. The F. malloryi leaves are the earliest appearance in the fossil record of a member of the witchalder genus Fothergilla, which includes the living species F. gardenii, and F. major, both of which are native to the southeastern United States. The genus also includes three or four other fossil species with two Asian Miocene species, F. viburnifolia from China, F. ryozenensis from Japan along with one Miocene North American species, F. praeolata of Oregon. Fothergilla durhamensis described from Eocene sediments in King County, Washington is considered dubious in placement, and it was transferred to the genus Platimeliphyllum by Huegele et al. (2021).
Corylopsis reedae is an extinct species of flowering plant in the family Hamamelidaceae known from fossil leaves found in the early Eocene Klondike Mountain Formation deposits of northern Washington state. C. readae is one of the oldest occurrences of the winter-hazel genus Corylopsis, which includes between seven and thirty species, all found in Asia. Fossils from two other occurrences are of similar age to C. readae, with Paleocene specimens from Greenland being placed in the form taxon Corylopsiphyllum and an Eocene Alaskan fossil being included in Corylopsis without species placement.
Stonebergia is an extinct genus in the rose family, Rosaceae, which contains the single species Stonebergia columbiana. The genus was described from a series of isolated fossil leaves in shale from an early Eocene location in southern British Columbia.
Langeria is an extinct genus of flowering plants in the family Hamamelidaceae, commonly known as the witch-hazel family, containing the solitary species Langeria magnifica. The species is known from fossil leaves found in the early Eocene deposits of northern Washington state, United States and similar aged formations in British Columbia, Canada.
Eoprephasma is an extinct genus of stick insect in the susumaniid subfamily Susumaniinae known from a group of Eocene fossils found in North America. When first described there was a single named species, Eoprephasma hichensi.
Eorpidae is a small family of extinct insects in the scorpionfly order, Mecoptera, which contains a single genus, Eorpa. Three Eocene age species found in Western North America have been placed into the genus: E. elverumi, E. jurgeni, and E. ypsipeda.
Tsukada is an extinct genus of flowering plant in the family Nyssaceae related to the modern "dove-tree", Davidia involucrata, containing the single species Tsukada davidiifolia. The genus is known from fossil leaves found in the early Eocene deposits of northern Washington state, United States and a similar aged formation in British Columbia, Canada.
Ulmus okanaganensis is an extinct species of flowering plant in the family Ulmaceae related to the modern elms. The species is known from fossil leaves, flowers, and fruits found in the early Eocene deposits of northern Washington state, United States and similar aged formations in British Columbia, Canada.
Betula leopoldae is an extinct species of birch in the family Betulaceae. The species is known from fossil leaves, catkins, and inflorescences found in the early Eocene deposits of northern Washington state, United States, and similar aged formations in British Columbia, Canada. The species is placed as basal in Betula, either as a stem group species, or an early divergent species.
Tetracentron hopkinsii is an extinct species of flowering plant in the family Trochodendraceae. The species is known from fossil leaves found in the early Eocene deposits of northern Washington state, United States and south Central British Columbia. The species was first described from fossil leaves found in the Allenby Formation. T. hopkinsii are possibly the leaves belonging to the extinct trochodendraceous fruits Pentacentron sternhartae.
Paraconcavistylon is an extinct genus of flowering plant in the family Trochodendraceae comprises a single species, Paraconcavistylon wehrii. The genus is known from fossil fruits and leaves found in the early Eocene deposits of northern Washington state, United States, and southern British Columbia, Canada. The species was initially described as a member of the related extinct genus Concavistylon as "Concavistylon" wehrii, but subsequently moved to the new genus Paraconcavistylon in 2020 after additional study.
Comptonia columbiana is an extinct species of sweet fern in the flowering plant family Myricaceae. The species is known from fossil leaves found in the early Eocene deposits of central to southern British Columbia, Canada, plus northern Washington state, United States, and, tentatively, the late Eocene of Southern Idaho and Earliest Oligocene of Oregon, United States.
Fagus langevinii is an extinct species of beech in the family Fagaceae. The species is known from fossil fruits, nuts, pollen, and leaves found in the early Eocene deposits of South central British Columbia, and northern Washington state, United States.