Gonyleptidae

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Gonyleptidae
Metagonyleptes calcar - 01.jpg
Metagonyleptes calcar Roewer, 1913, from Brazil
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Subphylum: Chelicerata
Class: Arachnida
Order: Opiliones
Superfamily: Gonyleptoidea
Family: Gonyleptidae
Sundevall, 1833
Subfamilies

See text

Gonyleptidae is a neotropical family of harvestmen (order Opiliones) with more than 800 species, the largest in the suborder Laniatores and the second largest of the Opiliones as a whole. The largest known harvestmen are gonyleptids.

Contents

Like most harvestmen, gonyleptids are almost exclusively nocturnal, except some Caelopyginae, Goniosomatinae (during reproductive season), Gonyleptinae, Mitobatinae, Pachylinae and Progonyleptoidellinae. Most species inhabit dense tropical, subtropical and temperate (Chile) forests, but some occur in open vegetation as the Pampas, the Cerrado, and the Caatinga. There are some species that live in caves, but only three troglobites are recorded for the family.

Name

The family is named after the type genus Gonyleptes, which is derived from Greek gony, gonatos = joint, knee + leptos, ê, on = thin, fine, delicate.

Diagnosis

Laniatores with coxa IV immensely developed, widely surpassing dorsal scutum in dorsal view in most species. Many species with double ozopore. Pedipalpus with cylindrical segments, strongly spined, tibia and tarsus flattened ventrally. Basal segments of leg IV with strong sexual dimorphism, shown either in spination, curvature or length. Penis with ventral plate well defined, glans may bear ventral and/or dorsal processes.

Distribution

Gonyleptidae have been recorded continuously from the southernmost tip of the South American continent (southern Chile and Argentina), Falklands to Costa Rica, with one isolated species cited from Guatemala.

Subfamilies

Gonyleptidae comprises several subfamilies, around 280 genera and over 700 species (early 2024):

Note: Metasarcinae was formerly included as a subfamily of Gonyleptidae and has since been raised to family level (Metasarcidae).

Relationships

Gonyleptidae is the sister-group of Cosmetidae and both are related to the Stygnidae and Cranaidae (Kury, 1992).

Related Research Articles

<span class="mw-page-title-main">Opiliones</span> Order of arachnids (harvestmen/daddy longlegs)

The Opiliones are an order of arachnids colloquially known as harvestmen, harvesters, harvest spiders, or daddy longlegs. As of April 2017, over 6,650 species of harvestmen have been discovered worldwide, although the total number of extant species may exceed 10,000. The order Opiliones includes five suborders: Cyphophthalmi, Eupnoi, Dyspnoi, Laniatores, and Tetrophthalmi, which were named in 2014.

<span class="mw-page-title-main">Laniatores</span> Suborder of harvestmen/daddy longlegs

Laniatores is the largest suborder of the arachnid order Opiliones with over 4,000 described species worldwide. The majority of the species are highly dependent on humid environments and usually correlated with tropical and temperate forest habitats.

<i>Lacronia</i> Genus of harvestmen/daddy longlegs

Lacronia is a genus of South American harvestmen, which includes four Brazilian species. A striking diagnostic character is the trochanter IV of male with strong medial prolateral apophysis forming a pincer with the dorso-apical apophysis of coxa IV.

<span class="mw-page-title-main">Pachylinae</span> Subfamily of harvestmen/daddy longlegs

Pachylinae is the most diverse subfamily of the harvestman family Gonyleptidae, including around 400 valid species. Major groups of species occur in the Brazilian Atlantic forest, Bolivian/Peruvian highlands, Argentina and Chilean temperate forest.

<span class="mw-page-title-main">Cosmetidae</span> Family of harvestmen/daddy longlegs

Cosmetidae is a family of harvestmen in the suborder Laniatores. With over 700 species, it is one of the largest families in Opiliones. They are endemic of the New World with a Nearctic-Neotropical distribution where a large fraction of the diversity of Opiliones are represented by this single family. Cosmetidae have the northern extent of their range into the USA, where a small number species occur in the southern states. However, the family is especially diverse in Mexico, Central America and northern South America; especially the Andean realms. Their range also extends further south into Argentina and southern Brazil, but they are absent in Chile. Cosmetidae are prevalent in Amazonian region, but only relatively few also occur in Brazilian Atlantic Forest. Several species are also found in the Caribbean.

<span class="mw-page-title-main">Cranaidae</span> Family of harvestmen/daddy longlegs

The Cranaidae are a family of neotropical harvestmen within the suborder Laniatores.

The Icaleptidae are a small family of neotropical harvestmen within the suborder Laniatores. Although only two species have been described, many more are probably to be discovered.

<span class="mw-page-title-main">Harvestman phylogeny</span> Order of arachnids

Harvestmen (Opiliones) are an order of arachnids often confused with spiders, though the two orders are not closely related. Research on harvestman phylogeny is in a state of flux. While some families are clearly monophyletic, that is share a common ancestor, others are not, and the relationships between families are often not well understood.

Caelopyginae is a neotropical sub-family of harvestmen in the family Gonyleptidae.

<i>Theromaster</i> Genus of harvestmen/daddy longlegs

Theromaster is a genus of armoured harvestmen in the family Cladonychiidae. There are at least two described species in Theromaster, found in the eastern United States.

<span class="mw-page-title-main">Travunioidea</span> Superfamily of harvestmen/daddy longlegs

Travunioidea is a superfamily of armoured harvestmen in the order Opiliones. There are 4 families and more than 70 described species in Travunioidea.

Wespus is a genus of armoured harvestmen in the family Phalangodidae. There is at least one described species in Wespus, W. arkansasensis.

<i>Vonones</i> (harvestman) Genus of harvestmen/daddy longlegs

Vonones is a genus of armoured harvestmen in the family Cosmetidae. There are at least two described species in Vonones.

Enigmina is a genus of armoured harvestmen in the family Phalangodidae. There are at least two described species in Enigmina.

<i>Crosbyella</i> Genus of harvestmen/daddy longlegs

Crosbyella is a genus of armoured harvestmen in the family Phalangodidae. There are about five described species in Crosbyella.

<i>Megacina</i> Genus of harvestmen/daddy longlegs

Megacina is a genus of armoured harvestmen in the family Phalangodidae. There are at least four described species in Megacina.

<i>Phalangodes</i> Genus of harvestmen/daddy longlegs

Phalangodes is a genus of armoured harvestmen in the family Phalangodidae. There is at least one described species in Phalangodes, P. armata.

<span class="mw-page-title-main">Cryptomastridae</span> Family of harvestmen/daddy longlegs

Cryptomastridae is a family of armoured harvestmen in the order Opiliones. There are two genera and four described species in Cryptomastridae, found in Oregon and Idaho.

Buemarinoa is a genus of armoured harvestmen in the family Cladonychiidae. There is one described species in Buemarinoa, B. patrizii, found in Sardinia, Italy.

Metasarcidae is a family of harvestmen, first described by Adriano B. Kury in 1994.

References