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Hymenoptera is a large order of insects, comprising the sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are parasitic. Females typically have a special ovipositor for inserting eggs into hosts or places that are otherwise inaccessible. This ovipositor is often modified into a stinger. The young develop through holometabolism —that is, they have a wormlike larval stage and an inactive pupal stage before they reach adulthood.
In evolutionary ecology, a parasitoid is an organism that lives in close association with its host at the host's expense, eventually resulting in the death of the host. Parasitoidism is one of six major evolutionary strategies within parasitism, distinguished by the fatal prognosis for the host, which makes the strategy close to predation.
The Scoliidae, the scoliid wasps, are a family of wasps comprising about 560 species worldwide. They tend to be black, often marked with yellow or orange, and their wing tips are distinctively corrugated. Males are more slender and elongated than females, with significantly longer antennae, but the sexual dimorphism is not as apparent as in many of the Tiphiidae and Thynnidae.
Sawflies are wasp-like insects that are in the suborder Symphyta within the order Hymenoptera, alongside ants, bees, and wasps. The common name comes from the saw-like appearance of the ovipositor, which the females use to cut into the plants where they lay their eggs. The name is associated especially with the Tenthredinoidea, by far the largest superfamily in the suborder, with about 7,000 known species; in the entire suborder, there are 8,000 described species in more than 800 genera. Symphyta is paraphyletic, consisting of several basal groups within the order Hymenoptera, each one rooted inside the previous group, ending with the Apocrita which are not sawflies.
The Ichneumonidae, also known as ichneumon wasps, ichneumonid wasps, ichneumonids, or Darwin wasps, are a family of parasitoid wasps of the insect order Hymenoptera. They are one of the most diverse groups within the Hymenoptera with roughly 25,000 species described as of 2016. However, this likely represents less than a quarter of their true richness as reliable estimates are lacking, along with much of the most basic knowledge about their ecology, distribution, and evolution. It is estimated that there are more species in this family than there are species of birds and mammals combined. Ichneumonid wasps, with very few exceptions, attack the immature stages of holometabolous insects and spiders, eventually killing their hosts. They thus fulfill an important role as regulators of insect populations, both in natural and semi-natural systems, making them promising agents for biological control.
Wasps in the family Pompilidae are commonly called spider wasps, spider-hunting wasps, or pompilid wasps. The family is cosmopolitan, with some 5,000 species in six subfamilies. Nearly all species are solitary, and most capture and paralyze prey, though members of the subfamily Ceropalinae are kleptoparasites of other pompilids, or ectoparasitoids of living spiders.
Parasitoid wasps are a large group of hymenopteran superfamilies, with all but the wood wasps (Orussoidea) being in the wasp-waisted Apocrita. As parasitoids, they lay their eggs on or in the bodies of other arthropods, sooner or later causing the death of these hosts. Different species specialise in hosts from different insect orders, most often Lepidoptera, though some select beetles, flies, or bugs; the spider wasps (Pompilidae) exclusively attack spiders.
Trigonalidae is a family of parasitic wasps in the suborder Apocrita. They are the only living members of the superfamily Trigonaloidea. Trigonalidae are divided into 2 subfamilies; Orthogonalinae and Trigonalinae. These wasps are extremely rare, but surprisingly diverse, with over 90 species in 16 genera, and are known from all parts of the world. It is possibly the sister group to all Aculeata.
Megalyroidea is a small hymenopteran superfamily of wasps that includes a single family, Megalyridae, with eight extant genera and 49 described species. Modern megalyrids are found primarily in the southern hemisphere, though fossils have only been found in the northern hemisphere. The most abundant and species-rich megalyrid fauna is in Australia. Another peak of diversity appears to be in the relict forests of Madagascar, but most of these species are still undescribed.
A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder. The wasps do not constitute a clade, a complete natural group with a single ancestor, as bees and ants are deeply nested within the wasps, having evolved from wasp ancestors. Wasps that are members of the clade Aculeata can sting their prey.
Maimetsha arctica is an extinct genus of wasp which existed during the Cretaceous period, and the only species in the genus Maimetsha.
Cretogonalys is an extinct genus of wasp which existed in Siberia during the late Cretaceous period, containing a single species, Cretogonalys taimyricus. It was described by Rasnitsyn in 1977.
Afrapia is an extinct genus of wasp which existed in Botswana during the late Cretaceous period. It contains the species Afrapia globularis and Afrapia variicornis.
Afromaimetsha robusta is an extinct species of wasp which existed in Botswana during the late Cretaceous period, and the only species in the genus Afromaimetsha.
Andyrossia is an extinct genus of wasp known from the Late Cretaceous Weald Clay of southern England, containing a single species, Andyrossia joyceae. It was first named by Rasnitsyn and Jarzembowski in 1998 as Arossia; this was later realised to be a junior homonym of a barnacle subgenus containing Concavus panamensis, and was replaced by the name Andyrossia in 2000.
Aptenoperissus is a genus of extinct wasp with eight described species, placed into the monotypic family Aptenoperissidae. The type species Aptenoperissus burmanicus resembles a mix between a grasshopper, an ant, and a wasp. It was described by a group of researchers from Oregon State University in a paper released online in October 2016. The piece of 100 million year old Burmese amber that it was preserved in was found in the Hukawng Valley of Myanmar in Southern Asia. A new family, Aptenoperissidae, was described to accommodate this insect. Species of Aptenoperissus were wingless, with a strong stinger. The creature had long legs making it capable of jumping higher than most insects. Subsequently additional species were described from the Myanmar amber: A. amabilis, A. delicatus, A. formosus, A. etius, A. magnifemoris, A. pusillus and A. zonalis. Initially placed in Ceraphronoidea, later studies placed it in Stephanoidea.
Trogus is a genus of parasitoid wasp found in the Holarctic and Neotropic regions. It is placed in the subfamily Ichneumoninae and the tribe Ichneumonini. Trogus species are parasites of larvae and pupae of the swallowtail butterfly family, Papilionidae. The genus consists of twelve extant and one extinct species.
2015 in paleoentomology is a list of new fossil insect taxa that were described during the year 2016, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.
Jouault, Corentin; Rosse-Guillevic, Simon. "A new genus of praeaulacid wasp from the mid-Cretaceous Kachin amber (Myanmar)". Annales de Paléontologie. 109 (1): 102599. doi:10.1016/j.annpal.2023.102599.
This paleoentomology list records new fossil insect taxa that were described during the year 2014, as well as notes other significant paleoentomology discoveries and events which occurred during that year.
References
- ↑ Vincent Perrichot, André Nel & D. Néraudeau (2004). "A new, enigmatic, evaniomorphan wasp in the Albian amber of France (Insecta: Hymenoptera)". Journal of Systematic Palaeontology . 2 (2): 159–162. Bibcode:2004JSPal...2..159P. doi:10.1017/S1477201904001245. S2CID 84325949.
- ↑ Alexandr P. Rasnitsyn & Denis J. Brothers (2009). "New genera and species of Maimetshidae (Hymenoptera: Stephanoidea s.l.) from the Turonian of Botswana, with comments on the status of the family" (PDF). African Invertebrates . 50 (1): 191–204. Bibcode:2009AfrIn..50..191R. doi: 10.5733/afin.050.0108 .
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