Herpetogaster Temporal range: Early to mid Cambrian: | |
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Specimens and diagram | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Superphylum: | Deuterostomia |
Stem group: | Ambulacraria |
Clade: | † Cambroernida |
Genus: | † Herpetogaster Caron et al., 2010 |
Species | |
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Herpetogaster is an extinct cambroernid genus of animal from the Early Cambrian Chengjiang biota of China, Blang Formation of China, Pioche Formation of Nevada and Middle Cambrian Burgess Shale of Canada containing the species Herpetogaster collinsi and Herpetogaster haiyanensis. [1] [2] [3] [4]
H. collinsi is known from over 160 specimens. It possessed a pair of branching tentacles and a tough but flexible body that curved helically to the right like a ram's horn and was divided into at least 13 segments. A flexible, extensible stolon emerged from the body at about the ninth segment and secured the animal to the sea floor, often by attaching to the sponge Vauxia. It is not known whether the attachment was permanent. [5]
A mouth opened between the tentacles, leading internally to a pharynx, a large lentil-shaped stomach, a narrower straight intestine, and an anus at the end of the "tail." The tentacles were softer than the body and probably extensible. A dark line running down the center of each tentacle and connecting with the head is tentatively reconstructed as a hydrostatic canal and/or vascular system; if this interpretation is correct, the tentacles might have been controlled by fluid pressure, and individuals could have fed by snaring either small prey or edible particles in the tentacles and bringing the tentacles to the mouth, as in living sea cucumbers. Structures on the back of the head have been noted as potential pharyngeal pores, which suggests a relationship with early echinoderms. The whole animal was 3-4 cm long. They were probably gregarious, as up to eight individuals have been found on the same slab. [1] The six specimens of Herpetogaster are known from the Greater Phyllopod bed, where they comprise less than 0.01% of the community. [6]
Hallucigenia is a genus of lobopodian known from Cambrian aged fossils in Burgess Shale-type deposits in Canada and China, and from isolated spines around the world. The generic name reflects the type species' unusual appearance and eccentric history of study; when it was erected as a genus, H. sparsa was reconstructed as an enigmatic animal upside down and back to front. Lobopodians are a grade of Paleozoic panarthropods from which the velvet worms, water bears, and arthropods arose.
The Maotianshan Shales (帽天山页岩) are a series of Early Cambrian sedimentary deposits in the Chiungchussu Formation, famous for their Konservat Lagerstätten, deposits known for the exceptional preservation of fossilized organisms or traces. The Maotianshan Shales form one of some forty Cambrian fossil locations worldwide exhibiting exquisite preservation of rarely preserved, non-mineralized soft tissue, comparable to the fossils of the Burgess Shale of British Columbia, Canada. They take their name from Maotianshan Hill in Chengjiang County, Yunnan Province, China.
Nectocaris is a genus of squid-like animal of controversial affinities known from the Cambrian period. The initial fossils were described from the Burgess Shale of Canada. Other similar remains possibly referrable to the genus are known from the Emu Bay Shale of Australia and Chengjiang Biota of China.
Sidneyia is an extinct arthropod known from fossils found from the Early to the Mid Cambrian of China and the Mid Cambrian Burgess Shale of British Columbia, Canada.
Ottoia is a stem-group archaeopriapulid worm known from Cambrian fossils. Although priapulid-like worms from various Cambrian deposits are often referred to Ottoia on spurious grounds, the only clear Ottoia macrofossils come from the Burgess Shale of British Columbia, which was deposited 508 million years ago. Microfossils extend the record of Ottoia throughout the Western Canada Sedimentary Basin, from the mid- to late- Cambrian. A few fossil finds are also known from China.
Canadaspis is an extinct genus of bivalved Cambrian arthropod, known from North America and China. They are thought to have been benthic feeders that moved mainly by walking and possibly used its biramous appendages to stir mud in search of food. They have been placed within the Hymenocarina, which includes other bivalved Cambrian arthropods.
Waptia is an extinct genus of arthropod from the Middle Cambrian of North America. It grew to a length of 6.65 cm (3 in), and had a large bivalved carapace and a segmented body terminating into a pair of tail flaps. It was an active swimmer and likely a predator of soft-bodied prey. It is also one of the oldest animals with direct evidence of brood care. Waptia fieldensis is the only species classified under the genus Waptia, and is known from the Burgess Shale Lagerstätte of British Columbia, Canada. Specimens of Waptia are also known from the Spence Shale of Utah, United States.
Dinomischusis an extinct genus of stalked filter-feeding animals within the Cambrian period, with specimens known from the Burgess Shale and the Maotianshan Shales. While long of uncertain affinities, recent studies have suggested it to be a stem-group ctenophore.
Leanchoilia is a megacheiran arthropod known from Cambrian deposits of the Burgess Shale in Canada and the Chengjiang biota of China.
Odaraia is a genus of bivalved arthropod from the Middle Cambrian. Its fossils, which reach 15 centimetres (5.9 in) in length, have been found in the Burgess Shale in British Columbia, Canada.
Branchiocaris is an extinct genus of Cambrian bivalved arthropod. The type and best known species, Branchiocaris pretiosa, was described from the Burgess Shale of British Columbia, Canada, in 1929, originally placed in Protocaris, and was placed into its own distinct genus by Briggs in 1976. Several other possible species have been described from Cambrian deposits in China, and it is also possibly known from Cambrian deposits in Utah. Branchiocaris pretiosa is around 80–90 millimetres (3.1–3.5 in) in length, with a highly segmented trunk, consisting of at least 44 ring-like segments, terminating in a forked tail telson. At the front of the animal is a pair of short segmented tapered antennules with at least 20 segments, as well as a pair of claw appendages. It was likely an active swimmer, and used the claw appendages to bring food to the mouth.
Alalcomenaeus is one of the most widespread and longest-surviving arthropod genera of the Early and Middle Cambrian. Known from over 300 specimens in the Burgess Shale and the Chengjiang biota. It is a member of the family Leanchoiliidae in the group Megacheira.
Plenocaris plena is a genus of extinct bivalved hymenocarine arthropod that lived in the Cambrian aged Burgess Shale and Chengjiang. Originally described as a species of Yohoia by Walcott in 1912, it was placed into its own genus in 1974.
Liangshanella is a genus of Cambrian bradoriid known from the Chengjiang biota and Burgess Shale. 6263 specimens of Liangshanella are known from the Greater Phyllopod bed, where they comprise 11.9% of the community.
Primicaris is genus of Cambrian arthropod from the Chengjiang biota of China and the Burgess Shale of Canada. It contains a single described species, P. larvaformis.
Eldonia is an extinct soft-bodied cambroernid animal of unknown affinity, best known from the Fossil Ridge outcrops of the Burgess Shale, particularly in the 'Great Eldonia layer' in the Walcott Quarry. In addition to the 550 collected by Walcott, 224 specimens of Eldonia are known from the Greater Phyllopod bed, where they comprise 0.43% of the community. Species also occur in the Chengjiang biota, Siberia, and in Upper Ordovician strata of Morocco.
Cambrorhytium is an enigmatic fossil genus known from the Latham Shale (California), and the Chengjiang (China) and Burgess Shale lagerstätte. 350 specimens of Cambrorhytium are known from the Greater Phyllopod bed, where they comprise 0.7% of the community.
Mollisonia is an extinct genus of Cambrian arthropod. Species are known from the Burgess Shale, Langston Formation, and Wheeler Shale of North America, as well as the Chengjiang Biota of China. Twenty-one specimens of Mollisonia are known from the Greater Phyllopod bed, where they comprise less than 0.1% of the community. Remains possibly attributable to the genus are also known from the Ordovician Fezouata Formation of Morocco and Bøggild Fjord Formation in Greenland. An observation published in 2019 suggests this genus is a basal chelicerate, closer to crown group Chelicerata than members of Habeliida. It is suggested to be closely related to Corcorania, Urokodia, and Thelxiope, which together form the clade Mollisoniida, which are thought to be closely related to Chelicerata.
Iotuba chengjiangensis is a 515 million year old Cambrian worm known from the Chengjiang biota. Originally interpreted as a phoronid, the organism is now recognized as an annelid cage worm affiliated with the Flabelligeridae and Acrocirridae, which Zhang et al grouped together in the new superfamily Flabelligeroidea.
Acinocricus is a genus of extinct panarthropod belonging to the group Lobopodia and known from the middle Cambrian Spence Shale of Utah, United States. As a monotypic genus, it has one species Acinocricus stichus. The only lobopodian discovered from the Spence Shale, it was described by Simon Conway Morris and Richard A. Robison in 1988. Owing to the original fragmentary fossils discovered since 1982, it was initially classified as an alga, but later realised to be an animal belonging to Cambrian fauna.