Lichenalia Temporal range: | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Bryozoa |
Class: | Stenolaemata |
Order: | † Cystoporida |
Family: | † Rhinoporidae |
Genus: | † Lichenalia |
Species [1] | |
See text |
Lichenalia is an extinct genus of cystoporate bryozoan belonging to the family Rhinoporidae. It is known from the Upper Ordovician to the Middle Silurian periods, which spanned from approximately 460 to 430 million years ago. The genus had a cosmopolitan distribution, with fossil specimens found in various regions of the world, including North America, Europe, and Asia.
The colonies of Lichenalia could either have a branched or tube-shaped form, or have an encrusting growth habit. The genus possessed prominent lunaria, which are shield-like structures that protected the zooids, or individual organisms, that made up the colony. The skeleton of Lichenalia was vesicular, meaning that it had a porous texture filled with numerous small chambers. The vesicular skeleton contained tunnels that appeared like ridges on the surface of the colony. The purpose of these tunnels is unknown, but they may have served as brooding chambers for the zooids.
Lichenalia was first described by the American paleontologist Edward Oscar Ulrich in 1882, based on fossil specimens collected from the Upper Ordovician rocks of Ohio. The genus was originally classified in the family Fistuliporidae, but subsequent studies have placed it in the family Rhinoporidae.
The ecology of Lichenalia is not well understood, but the genus is believed to have been a filter-feeding organism that lived in shallow marine environments. Like other bryozoans, Lichenalia colonies were composed of many individual zooids that were interconnected by tiny tubes. The zooids fed on microscopic organisms that they captured from the surrounding water using a ring of ciliated tentacles called a lophophore. As filter-feeders, bryozoans such as Lichenalia played a role in controlling the population of plankton in ancient marine ecosystems.
Lichenalia colonies could form hollow branched or tube-shaped colonies, or have an encrusting growth habit. The encrusting growth form is particularly common in rocks of the Middle Silurian period. Some species of Lichenalia have been found in association with other reef-building organisms, such as corals and stromatoporoids. This suggests that Lichenalia may have played a role in building or stabilizing reef structures.
Lichenalia belongs to the class Stenolaemata within the phylum Bryozoa. The genus is classified in the family Rhinoporidae, which is characterized by a vesicular skeleton with tunnels that appear as ridges on the surface of the colony. Other genera in the Rhinoporidae include Rhinopora, Fistulipora, and Heterotrypa.
Several species of Lichenalia have been described, including:
Fossil specimens of Lichenalia have been found in various parts of the world, including North America (e.g., Ohio, Tennessee, and New York), Europe (e.g., Sweden and the Czech Republic), and Asia (e.g., China and Kazakhstan). The genus has been recovered from rocks of Upper Ordovician to Middle Silurian age, indicating that it existed for a period of approximately 30 million years.
The ecology of Lichenalia is poorly understood, as there is limited information available on its biology and behavior. Like other bryozoans, Lichenalia was a filter-feeding organism that likely lived in shallow marine environments. The genus may have played a role in building or stabilizing reef structures, as some species have been found in association with other reef-building organisms, such as corals and stromatoporoids.
Bryozoa are a phylum of simple, aquatic invertebrate animals, nearly all living in sedentary colonies. Typically about 0.5 millimetres long, they have a special feeding structure called a lophophore, a "crown" of tentacles used for filter feeding. Most marine bryozoans live in tropical waters, but a few are found in oceanic trenches and polar waters. The bryozoans are classified as the marine bryozoans (Stenolaemata), freshwater bryozoans (Phylactolaemata), and mostly-marine bryozoans (Gymnolaemata), a few members of which prefer brackish water. 5,869 living species are known. At least two genera are solitary ; the rest are colonial.
Graptolites are a group of colonial animals, members of the subclass Graptolithina within the class Pterobranchia. These filter-feeding organisms are known chiefly from fossils found from the Middle Cambrian through the Lower Carboniferous (Mississippian). A possible early graptolite, Chaunograptus, is known from the Middle Cambrian. Recent analyses have favored the idea that the living pterobranch Rhabdopleura represents an extant graptolite which diverged from the rest of the group in the Cambrian. Fossil graptolites and Rhabdopleura share a colony structure of interconnected zooids housed in organic tubes (theca) which have a basic structure of stacked half-rings (fuselli). Most extinct graptolites belong to two major orders: the bush-like sessile Dendroidea and the planktonic, free-floating Graptoloidea. These orders most likely evolved from encrusting pterobranchs similar to Rhabdopleura. Due to their widespread abundance, planktonic lifestyle, and well-traced evolutionary trends, graptoloids in particular are useful index fossils for the Ordovician and Silurian periods.
Entoprocta, or Kamptozoa, is a phylum of mostly sessile aquatic animals, ranging from 0.1 to 7 millimetres long. Mature individuals are goblet-shaped, on relatively long stalks. They have a "crown" of solid tentacles whose cilia generate water currents that draw food particles towards the mouth, and both the mouth and anus lie inside the "crown". The superficially similar Bryozoa (Ectoprocta) have the anus outside a "crown" of hollow tentacles. Most families of entoprocts are colonial, and all but 2 of the 150 species are marine. A few solitary species can move slowly.
Stenolaemata are a class of exclusively marine bryozoans. Stenolaemates originated and diversified in the Ordovician, and more than 600 species are still alive today. All extant (living) species are in the order Cyclostomatida, the third-largest order of living bryozoans.
Membranipora membranacea is a very widely distributed species of marine bryozoan known from the Atlantic and Pacific Oceans, usually in temperate zone environments. This bryozoan is a colonial organism characterized by a thin, mat-like encrustation, white to gray in color. It may be known colloquially as the coffin box, sea-mat or lacy crust bryozoan and is often abundantly found encrusting seaweeds, particularly kelps.
Bioerosion describes the breakdown of hard ocean substrates – and less often terrestrial substrates – by living organisms. Marine bioerosion can be caused by mollusks, polychaete worms, phoronids, sponges, crustaceans, echinoids, and fish; it can occur on coastlines, on coral reefs, and on ships; its mechanisms include biotic boring, drilling, rasping, and scraping. On dry land, bioerosion is typically performed by pioneer plants or plant-like organisms such as lichen, and mostly chemical or mechanical in nature.
Cyclostomatida, or cyclostomata, are an ancient order of stenolaemate bryozoans which first appeared in the Lower Ordovician. It consists of 7+ suborders, 59+ families, 373+ genera, and 666+ species. The cyclostome bryozoans were dominant in the Mesozoic; since that era, they have decreased. Currently, cyclostomes seldom constitute more than 20% of the species recorded in regional bryozoan faunas.
Stromatoporoidea is an extinct clade of sea sponges common in the fossil record from the Middle Ordovician to the Late Devonian. They can be characterized by their densely layered calcite skeletons lacking spicules. Stromatoporoids were among the most abundant and important reef-builders of their time, living close together in flat biostromes or elevated bioherms on soft tropical carbonate platforms.
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Constellaria is an extinct genus of bryozoan from the Middle Ordovician to Early Silurian from North America, Asia and Europe. These branching coral-like bryozoans formed bushy colonies 10-15 mm across on the seabed. The fairly thick branches were erect, often compressed in one direction, and covered with distinctive tiny, star-shaped mounds called maculae or monticules. Feeding zoids were located along the rays of the stars. The maculae probably formed "chimneys" for the expulsion of exhalant feeding currents from the surface of a colony, after water had been filtered to obtain food for the organisms.
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Amathia verticillata, commonly known as the spaghetti bryozoan, is a species of colonial bryozoans with a bush-like structure. It is found in shallow temperate and warm waters in the western Atlantic Ocean and the Caribbean Sea and has spread worldwide as a fouling organism. It is regarded as an invasive species in some countries.
Fenestellidae is a family of bryozoans belonging to the order Fenestrida. The skeleton of its colonies consists of stiff branches that are interconnected by narrower crossbars. The individuals of the colony inhabit one side of the branches in two parallel rows or two at the branch base and three or more rows further up. Zooids can be recognized as small rimmed pores, and in well-preserved specimens the apertures are closed by centrally perforated lids. The front of the branches carries small nodes in a row or zigzag line between the apertures. Branches split from time to time giving the colonies a fan-shape or, in the genus Archimedes, create an mesh in the shape of an Archimedes screw.
Fenestella is a genus of bryozoans or moss animals, forming fan–shaped colonies with a netted appearance. It is known from the Middle Ordovician to the early Upper Triassic (Carnian), reaching its largest diversity during the Carboniferous. Many hundreds of species have been described from marine sediments all over the world.
Cryptosula pallasiana is a species of colonial bryozoan in the order Cheilostomatida. It is native to the Atlantic Ocean where it occurs in northwestern Europe and northern Africa, and the eastern seaboard of North America. It has been accidentally introduced to the western coast of North America and to other parts of the world.
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Beania magellanica is a species of colonial bryozoan in the family Beaniidae. It has a cosmopolitan distribution, occurring in shallow waters in the Atlantic and Pacific Oceans and in Antarctica.
Chorizopora brongniartii is a species of bryozoan in the family Chorizoporidae. It is an encrusting bryozoan, the colonies forming spreading patches. It has a widespread distribution in tropical and temperate seas.
Homotrypa is an extinct genus of bryozoans from the Ordovician and Silurian periods, known from fossils found in the United States. Its colonies are branch-like and have small monticules made of groups of three or four larger zooecia slightly protruding out from the main surface of the colony. In cross section, the zooecia are erect in axis and gently curve toward the surface of the colony.