Proboscidea is a taxonomic order of afrotherian mammals containing one living family (Elephantidae) and several extinct families. First described by J. Illiger in 1811, it encompasses the elephants and their close relatives. Proboscideans include some of the largest known land mammals. The largest land mammal of all time may have been a proboscidean; the elephant Palaeoloxodon namadicus has been estimated to be up to 5.2 m (17.1 ft) at the shoulder and may have weighed up to 22 t, surpassing the paraceratheres, the otherwise largest known land mammals, though this estimate was made based on a single fragmentary femur and is speculative. The largest extant proboscidean is the African bush elephant, with a record of size of 4 m (13.1 ft) at the shoulder and 10.4 t. In addition to their enormous size, later proboscideans are distinguished by tusks and long, muscular trunks, which were less developed or absent in early proboscideans.
A mastodon is any proboscidean belonging to the extinct genus Mammut. Mastodons inhabited North and Central America from the late Miocene up to their extinction at the end of the Pleistocene 10,000 to 11,000 years ago. Mastodons are the most recent members of the family Mammutidae, which diverged from the ancestors of elephants at least 25 million years ago. M. americanum, the American mastodon, is the youngest and best-known species of the genus. They lived in herds and were predominantly forest-dwelling animals. M. americanum is inferred to have had a browsing diet with a preference for woody material, distinct from that of the contemporary Columbian mammoth. Mastodons became extinct as part of the Quaternary extinction event that exterminated most Pleistocene megafauna present in the Americas, believed to have been caused by a combination of climate changes at the end of the Pleistocene and hunting by recently arrived Paleo-Indians, as evidenced by a number of kill sites where mastodon remains are associated with human artifacts.
Elephantidae is a family of large, herbivorous proboscidean mammals collectively called elephants and mammoths. These are large terrestrial mammals with a snout modified into a trunk and teeth modified into tusks. Most genera and species in the family are extinct. Only two genera, Loxodonta and Elephas, are living.
Elasmotherium is an extinct genus of large rhinoceros endemic to Eurasia during Late Miocene through to the Late Pleistocene, with the youngest reliable dates around 39,000 years ago. It was the last surviving member of Elasmotheriinae, a distinctive group of rhinoceroses separate from the group that contains living rhinoceros (Rhinocerotinae).
Elephas is one of two surviving genera in the family of elephants, Elephantidae, with one surviving species, the Asian elephant, Elephas maximus. Several extinct species have been identified as belonging to the genus, extending back to the Pliocene era.
Palaeoloxodon is an extinct genus of elephant. The genus originated in Africa during the Early Pleistocene, and expanded into Eurasia at the beginning of the Middle Pleistocene. The genus contains some of the largest known species of elephants, over 4 metres (13 ft) tall at the shoulders, including the African Palaeoloxodon recki, the European straight-tusked elephant and the South Asian Palaeoloxodon namadicus. P. namadicus has been suggested to be the largest known land mammal by some authors based on extrapolation from fragmentary remains, though these estimates are highly speculative. In contrast, the genus also contains many species of dwarf elephants that evolved via insular dwarfism on islands in the Mediterranean, some only 1 metre (3.3 ft) in height, making them the smallest elephants known. The genus has a long and complex taxonomic history, and at various times, it has been considered to belong to Loxodonta or Elephas, but today is usually considered a valid and separate genus in its own right.
Gomphotheres are an extinct group of proboscideans related to modern elephants. They were widespread across Afro-Eurasia and North America during the Miocene and Pliocene epochs and dispersed into South America during the Pleistocene as part of the Great American Interchange. Gomphotheres are a paraphyletic group that is ancestral to Elephantidae, which contains modern elephants, as well as Stegodontidae. While most famous forms such as Gomphotherium had long lower jaws with tusks, which is the ancestral condition for the group, some later members developed shortened (brevirostrine) lower jaws with either vestigial or no lower tusks, looking very similar to modern elephants, an example of parallel evolution, which outlasted the long-jawed gomphotheres. By the end of the Early Pleistocene, gomphotheres became extinct in Afro-Eurasia, with the last two genera, Cuvieronius ranging from southern North America to western South America, and Notiomastodon having a wide range over most of South America until the end of the Pleistocene around 12,000 years ago, when they became extinct following the arrival of humans.
Dwarf elephants are prehistoric members of the order Proboscidea which, through the process of allopatric speciation on islands, evolved much smaller body sizes in comparison with their immediate ancestors. Dwarf elephants are an example of insular dwarfism, the phenomenon whereby large terrestrial vertebrates that colonize islands evolve dwarf forms, a phenomenon attributed to adaptation to resource-poor environments and lack of predation and competition. Some modern populations of Asian elephants have also undergone size reduction on islands to a lesser degree, resulting in populations of pygmy elephants.
African elephants are members of the genus Loxodonta comprising two living elephant species, the African bush elephant and the smaller African forest elephant. Both are social herbivores with grey skin, but differ in the size and colour of their tusks and in the shape and size of their ears and skulls.
Antidorcas is a genus of antelope that includes the living springbok and several fossil species.
Palaeoloxodon recki, often known by the synonym Elephas recki is an extinct species of elephant native to Africa and West Asia from the Pliocene or Early Pleistocene to the Middle Pleistocene. During most of its existence, the species in its broadest sense represented the dominant elephant species in East Africa. While the type subspecies P. recki recki as well as P. recki ileretensis are widely accepted to be closely related to Eurasian Palaeoloxodon, the relationships of the other, chronologically earlier subspecies to P. recki recki and P. recki ileretensis are uncertain, with it being suggested they are unrelated and should be elevated to separate species.
The Columbian mammoth is an extinct species of mammoth that inhabited the Americas as far north as the Northern United States and as far south as Costa Rica during the Pleistocene epoch. The Columbian mammoth descended from Eurasian mammoths that colonised North America during the Early Pleistocene around 1.5-1.3 million years ago, and later experienced hybridisation with the woolly mammoth lineage. The Columbian mammoth was among the last mammoth species, and the pygmy mammoths evolved from them on the Channel Islands of California. The closest extant relative of the Columbian and other mammoths is the Asian elephant.
The straight-tusked elephant is an extinct species of elephant that inhabited Europe and Western Asia during the Middle and Late Pleistocene. Recovered individuals have reached up to 3.81–4.2 metres (12.5–13.8 ft) in shoulder height, and an estimated 11.3–15 tonnes in weight. Like modern elephants, the straight-tusked elephant lived in herds, flourishing during interglacial periods, when its range would extend as far north as Great Britain. Skeletons found in association with stone tools and wooden spears suggest they were scavenged and hunted by early humans, including Neanderthals. It is the ancestral species of most dwarf elephants that inhabited islands in the Mediterranean.
Anancus is an extinct genus of "tetralophodont gomphothere" native to Afro-Eurasia, that lived from the Tortonian stage of the late Miocene until its extinction during the Early Pleistocene, roughly from 8.5–2 million years ago.
Mammuthus trogontherii, sometimes called the steppe mammoth, is an extinct species of mammoth that ranged over most of northern Eurasia during the Early and Middle Pleistocene, approximately 1.7 million-200,000 years ago. One of the largest mammoth species, it evolved in East Asia during the Early Pleistocene, around 1.8 million years ago, before migrating into North America around 1.5 million years ago, and into Europe during the Early/Middle Pleistocene transition, around 1 to 0.7 million years ago. It was the ancestor of the woolly mammoth and Columbian mammoth of the later Pleistocene.
Pelorovis is an extinct genus of African wild cattle which existed during the Pleistocene epoch. The best known species is Pelorovis oldowayensis from Olduvai Gorge in Tanzania, from the Early Pleistocene. The species "Pelorovis" antiquus from the Late Pleistocene-Holocene has since been moved into Syncerus, the same genus as living African buffalo.
The Late Pleistocene to the beginning of the Holocene saw numerous extinctions of predominantly megafaunal (large) animal species, which resulted in a collapse in faunal density and diversity across the globe. The extinctions during the Late Pleistocene are differentiated from previous extinctions by the widespread absence of ecological succession to replace these extinct megafaunal species, and the regime shift of previously established faunal relationships and habitats as a consequence. The timing and severity of the extinctions varied by region and are thought to have been driven by varying combinations of human and climatic factors. Human impact on megafauna populations is thought to have been driven by hunting ("overkill"), as well as possibly environmental alteration. The relative importance of human vs climatic factors in the extinctions has been the subject of long-running controversy.
The African bush elephant, also known as the African savanna elephant, is one of two extant African elephant species and one of three extant elephant species. It is the largest living terrestrial animal, with bulls reaching a shoulder height of up to 3.96 m and a body mass of up to 10.4 t.
Loxodonta adaurora is an extinct species of elephant in the genus Loxodonta, that of the African elephants. Fossils of Loxodonta adaurora have only been found in Africa, where they developed in the Pliocene. L. adaurora was presumed to be the genetic antecedent of the two modern African elephant species; however, an analysis in 2009 suggested that L. africana evolved from L. atlantica. The same study concluded that Loxodonta adaurora was morphologically indistinguishable from Mammuthus subplanifrons and that these constituted the same species probably within the mammoth lineage However, other authors have continued to consider L. adaurora a valid species of Loxodonta, with some considering it an early morph of Loxodonta exoptata.
Syncerus antiquus is an extinct species of buffalo from the Late Pleistocene and Holocene of Africa. It was one of the largest species in its family, potentially weighing up to 2,000 kilograms (4,400 lb). Due to this fact, it is sometimes known as the African giant buffalo. The time of its extinction is of debate; Syncerus antiquus either became extinct at the end of the Late Pleistocene about 12,000 years ago or during the Holocene, some 4,000 years ago.
