Ungulates are members of the diverse clade Euungulata which primarily consists of large mammals with hooves. Once part of the clade "Ungulata" along with the clade Paenungulata, "Ungulata" has since been determined to be a polyphyletic and thereby invalid clade based on molecular data. As a result, true ungulates had since been reclassified to the newer clade Euungulata in 2001 within the clade Laurasiatheria while Paenungulata has been reclassified to a distant clade Afrotheria. Living ungulates are divided into two orders: Perissodactyla including equines, rhinoceroses, and tapirs; and Artiodactyla including cattle, antelope, pigs, giraffes, camels, sheep, deer, and hippopotamuses, among others. Cetaceans such as whales, dolphins, and porpoises are also classified as artiodactyls, although they do not have hooves. Most terrestrial ungulates use the hoofed tips of their toes to support their body weight while standing or moving. Two other orders of ungulates, Notoungulata and Litopterna, both native to South America, became extinct at the end of the Pleistocene, around 12,000 years ago.
Artiodactyls are mammals belonging to the order Artiodactyla. Typically, they are ungulates which bear weight equally on two of their five toes: the third and fourth, often in the form of a hoof. The other three toes are either present, absent, vestigial, or pointing posteriorly. By contrast, most perissodactyls bear weight on an odd number of the five toes. Another difference between the two is that many artiodactyls digest plant cellulose in one or more stomach chambers rather than in their intestine as perissodactyls do. The advent of molecular biology, along with new fossil discoveries, found that cetaceans fall within this taxonomic branch, being most closely related to hippopotamuses. Some modern taxonomists thus apply the name Cetartiodactyla to this group, while others opt to include cetaceans within the existing name of Artiodactyla. Some researchers use "even-toed ungulates" to exclude cetaceans and only include terrestrial artiodactyls, making the term paraphyletic in nature.
Horses can use various gaits during locomotion across solid ground, either naturally or as a result of specialized training by humans.
Camelids are members of the biological family Camelidae, the only currently living family in the suborder Tylopoda. The seven extant members of this group are: dromedary camels, Bactrian camels, wild Bactrian camels, llamas, alpacas, vicuñas, and guanacos. Camelids are even-toed ungulates classified in the order Artiodactyla, along with species including whales, pigs, deer, cattle, and antelopes.
Stenomylus is an extinct genus of miniature camelid native to North America that died out around 30 million years ago. Its name is derived from the Greek στενός and μύλος.
Procamelus is an extinct genus of camel endemic to North America. It lived from the Middle to Late Miocene 16.3—5.3 mya, existing for approximately 11 million years. The name is derived from the Greek πρό, meaning "before" or denoting priority of order, and κάμελος ("camel"), thus meaning "fore-camel", "early camel" or "predecessor camel".
Pedopenna is a genus of small, feathered, maniraptoran dinosaur from the Daohugou Beds in China. It is possibly older than Archaeopteryx, though the age of the Daohugou Beds where it was found is debated. A majority of studies suggest that beds probably date from between the late Middle Jurassic and early Late Jurassic Period.
Ornithosuchus is an extinct genus of pseudosuchians from the Late Triassic (Carnian) Lossiemouth Sandstone of Scotland. It was originally thought to be the ancestor to the carnosaurian dinosaurs, but it is now known to be more closely related to crocodilians than to dinosaurs.
Keraterpeton is an extinct genus of lepospondyl amphibian, previously included within the monotypic Keraterpedontidae family, from the Carboniferous period of Europe and North America ; it is the oldest known member of the family Diplocaulidae.
Thadeosaurus is an extinct genus of diapsid reptile belonging to the family Younginidae. Fossils have been found in the Lower Sakamena Formation of the Morondava Basin, Madagascar in 1981, and date to the late Permian to the early Triassic period.
Merycoidodon is an extinct genus of herbivorous artiodactyl of the family Merycoidodontidae, more popularly known by the name Oreodon. It was endemic to North America during the Middle Eocene to Middle Miocene existing for approximately 30 million years.
Homalodotherium is an extinct genus of notoungulate mammals native to South America. Fossils of Homalodotherium have been found in the Middle Miocene Santa Cruz Formation of Argentina and the Río Frías Formation of Chile.
Eotitanops is an extinct genus of brontothere native to North America and Asia.
Elomeryx is an extinct genus of artiodactyl ungulate, and is among the earliest known anthracotheres. The genus was extremely widespread, first being found in Asia in the middle Eocene, in Europe during the latest Eocene, and having spread to North America by the early Oligocene. The closest living relatives of the Elomeryx are bovids, suids, and cetaceans.
Poebrotherium is an extinct genus of camelid, endemic to North America. They lived from the Eocene to Miocene epochs, 46.3—13.6 mya, existing for approximately 32 million years.
Barylambda is an extinct genus of pantodont mammal from the middle to late Paleocene, well known from several finds in the Wasatchian DeBeque Formation of Colorado and the Clarkforkian Wasatch Formation to Tiffanian Fort Union Formation in Wyoming. Three species of Barylambda are currently recognized. The creature likely lived a life similar to that of a modern tapir, browsing on foliage and soft vegetation. Barylambda seems to have been quite successful for an early pantodont, though eventually it seems to have been replaced in its ecosystem by other pantodonts, such as Coryphodon.
The evolution of the horse, a mammal of the family Equidae, occurred over a geologic time scale of 50 million years, transforming the small, dog-sized, forest-dwelling Eohippus into the modern horse. Paleozoologists have been able to piece together a more complete outline of the evolutionary lineage of the modern horse than of any other animal. Much of this evolution took place in North America, where horses originated but became extinct about 10,000 years ago, before being reintroduced in the 15th century.
The limbs of the horse are structures made of dozens of bones, joints, muscles, tendons, and ligaments that support the weight of the equine body. They include two apparatuses: the suspensory apparatus, which carries much of the weight, prevents overextension of the joint and absorbs shock, and the stay apparatus, which locks major joints in the limbs, allowing horses to remain standing while relaxed or asleep. The limbs play a major part in the movement of the horse, with the legs performing the functions of absorbing impact, bearing weight, and providing thrust. In general, the majority of the weight is borne by the front legs, while the rear legs provide propulsion. The hooves are also important structures, providing support, traction and shock absorption, and containing structures that provide blood flow through the lower leg. As the horse developed as a cursorial animal, with a primary defense mechanism of running over hard ground, its legs evolved to the long, sturdy, light-weight, one-toed form seen today.
Isasicursor is a genus of elasmarian ornithopod from the Chorrillo Formation from Santa Cruz Province in Argentina. The type and only species is Isasicursor santacrucensis. It was a contemporary of the sauropod Nullotitan which was described in the same paper.
