A cactus is a member of the plant family Cactaceae, a family comprising about 127 genera with some 1,750 known species of the order Caryophyllales. The word cactus derives, through Latin, from the Ancient Greek word κάκτος (káktos), a name originally used by Theophrastus for a spiny plant whose identity is now not certain. Cacti occur in a wide range of shapes and sizes. They are native to the Americas, ranging from Patagonia in the south to parts of western Canada in the north, with the exception of Rhipsalis baccifera, which is also found in Africa and Sri Lanka. Cacti are adapted to live in very dry environments, including the Atacama Desert, one of the driest places on Earth. Because of this, cacti show many adaptations to conserve water. For example, almost all cacti are succulents, meaning they have thickened, fleshy parts adapted to store water. Unlike many other succulents, the stem is the only part of most cacti where this vital process takes place. Most species of cacti have lost true leaves, retaining only spines, which are highly modified leaves. As well as defending against herbivores, spines help prevent water loss by reducing air flow close to the cactus and providing some shade. In the absence of true leaves, cacti's enlarged stems carry out photosynthesis.
Echinopsis is a genus of cacti native to South America, sometimes known as hedgehog cactus, sea-urchin cactus or Easter lily cactus. As of October 2023, there are about 20 accepted species, ranging from large and treelike types to small globose cacti. The name derives from echinos hedgehog or sea urchin, and opsis appearance, a reference to these plants' dense coverings of spines.
Pereskia is a small genus of about four species of cacti that do not look much like other types of cacti, having substantial leaves and non-succulent stems. The genus is named after Nicolas-Claude Fabri de Peiresc, a 16th-century French botanist. The genus was more widely circumscribed until molecular phylogenetic studies showed that it was paraphyletic. The majority of species have since been transferred to Leuenbergeria and Rhodocactus. Although Pereskia does not resemble other cacti in its overall morphology, close examination shows spines developing from areoles, and the distinctive floral cup of the cactus family.
In 1984, the International Organization for Succulent Plant Study set up a working party, now called the International Cactaceae Systematics Group, to produce a consensus classification of the cactus family, down to the level of genus. Their classification has been used as the basis for systems published since the mid-1990s. Treatments in the 21st century have generally divided the family into around 125–130 genera and 1,400–1,500 species, which are then arranged in a number of tribes and subfamilies. However, subsequent molecular phylogenetic studies have shown that a very high proportion of the higher taxa are not monophyletic, i.e. they do not contain all of the descendants of a common ancestor. As of August 2023, the internal classification of the family Cactaceae remained uncertain and subject to change. A classification incorporating many of the insights from the molecular studies was produced by Nyffeler and Eggli in 2010.
Armatocereus is a genus of mostly tree-like cacti from South America. These species have a conspicuous constriction at the end of the annual growth. The flowers are mostly white, with a more or less spiny ovary. The fruits are mostly spiny.
Thelocactus is a genus of flowering plants in the cactus family Cactaceae. Members of the genus are native to the arid lands of Central and Northern Mexico.
Hatiora is a small genus of epiphytic cacti which belongs to the tribe Rhipsalideae within the subfamily Cactoideae of the Cactaceae. Recent taxonomic studies have led to the three species formerly placed in subgenus Rhipsalidopsis being removed from the genus, including the well known and widely cultivated ornamental plants known as Easter cactus or Whitsun cactus.
Soehrensia is a genus of cacti native to South America.
Vatricania is a monotypic genus of flowering plants belonging to the family Cactaceae. It contains just one species, Vatricania guentheri. It has the common name of red tail cactus.
Mila caespitosa is a species of cacti and the only species of the genus Mila. Its generic name is an anagram of Lima, Peru, the city near which the plant is found. The genus was first thought to comprise 13 species, until recent studies suggest they form one very variable species.
Weingartia is a genus in the family Cactaceae, with species native to Bolivia and Argentina. Molecular phylogenetic evidence suggests that it may be distinct from Rebutia. It is treated as a synonym of that genus by Plants of the World Online as of September 2023, but recognized as an alternative generic name in the third edition of the CITES Cactaceae Checklist. It may also be treated as Rebutia subg. Weingartia.
Pereskiopsis is a genus of cactus in the subfamily Opuntioideae. Unlike typical cacti, it has persistent fleshy leaves. The genus name refers to its resemblance to the genus Pereskia. Most species are found in Mexico south through Guatemala to Honduras, with one species in Bolivia. The incorrect spelling Peireskiopsis has also been used.
Rhodocactus is a genus of flowering plant in the cactus family Cactaceae, native to central South America. Unlike most species of cacti, Rhodocactus has persistent leaves and a fully tree-like habit. The genus was sunk into a broadly circumscribed Pereskia, but molecular phylogenetic studies from 2005 onwards showed that with this circumscription Pereskia was paraphyletic, and in 2016, Rhodocactus was restored for southern South American species.
Rhodocactus grandifolius is a species of cactus native to eastern and southern Brazil. Like all species in the genus Rhodocactus and unlike most cacti, it has persistent leaves. It was first described in 1819. It is grown as an ornamental plant and has naturalized outside its native range.
Rhodocactus bahiensis is a species of tree-like cactus that is endemic to the Brazilian state of Bahia. First described as Pereskia bahiensis, it was transferred to Rhodocactus in 2016. Like all species in the genus Rhodocactus, and unlike most cacti, it has persistent leaves. In its native locality, it is used to form hedges.
Rhodocactus stenanthus is a species of cactus that is endemic to Brazil. First described as Pereskia stenantha, it was transferred to Rhodocactus in 2016. Like all species in the genus Rhodocactus, and unlike most cacti, it has persistent leaves. In its native locality, it is sometimes used in hedges.
Lophocereus marginatus is a species of plant in the family Cactaceae. It is sometimes called Mexican fencepost cactus.
Pereskia aculeata is a scrambling shrub in the family Cactaceae. Common names include Barbados gooseberry, blade-apple cactus, leaf cactus, rose cactus, and lemonvine. It is native to tropical America. The leaves and fruits are edible, containing high quantities of protein, iron and other nutrients, and it is a popular vegetable in parts of the Brazilian state of Minas Gerais under the name of ora-pro-nóbis.
Leuenbergeria is a genus of flowering plant in the family Cactaceae, mostly native around the Caribbean. Unlike most cacti, it has persistent leaves and develops bark on its stems early in its growth. The genus was created in 2012 by Joël Lodé. Before the creation of Leuenbergeria as a genus, the species within it were included in a broadly circumscribed genus, Pereskia. Leuenbergeria is the only genus in the subfamily Leuenbergerioideae.
Rhodocactus nemorosus is a species of flowering plant in the cactus family Cactaceae, native to southern Brazil, Paraguay, Uruguay and northeast Argentina. Like all species in the genus Rhodocactus and unlike most cacti, it has persistent leaves. It was first described by Nicolás Rojas Acosta in 1897 as Pereskia nemorosa, and transferred to Rhodocactus in 2016.
