Schoenesmahl

Last updated

Schoenesmahl
Temporal range: Late Jurassic, 150.8  Ma
O
S
D
C
P
T
J
K
Pg
N
Compsognathus by Nopcsa, 1903.jpg
1903 illustration of the holotype by Franz Nopcsa
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Squamata
Family: Ardeosauridae
Genus: Schoenesmahl
Conrad, 2018
Species:
S. dyspepsia
Binomial name
Schoenesmahl dyspepsia
Conrad, 2018

Schoenesmahl is an extinct genus of lizard from the Late Jurassic Painten Formation of Germany. [1] It contains only a single species, S. dyspepsia. [2]

It is known only from specimen SNSB-BSPG AS I 563b, a single disarticulated specimen (consisting of an incomplete skeleton lacking nasals, vomers, palatines, postorbitals, quadrates, anterior presacral vertebrae, pectoral girdles, most of the radii and ulnae, manus, ilium, ischium, tarsals and the distal pedal phalanges form digits I, II, IV and V) [2] preserved in the stomach of the holotype specimen of the small theropod dinosaur Compsognathus longipes , which was discovered by Joseph Oberndorfer in Kelheim, Bavaria no later than 1859. [3] In 1866, Oberndorfer's collection, including the Schoenesmahl dyspepsia and Compsognathus longipes holotype specimens, was acquired by the paleontological state collection in Munich. [4] Othniel Charles Marsh, who examined the specimen in 1881, thought that this small skeleton in the Compsognathus belly was an embryo, but in 1903, Franz Nopcsa concluded that it was a lizard. [5] In 1978, John Ostrom identified the remains as belonging to a lizard of the genus Bavarisaurus , [6] which he concluded was a fast and agile runner owing to its long tail and limb proportions.

This specimen was long classified in the genus Bavarisaurus , [7] but a 2018 study found it to be a distinct taxon most closely related to Ardeosaurus and reclassified it as its own genus. The status of the specimen as prey for Compsognathus is reflected in the genus and species name, with Schoenesmahl deriving from schöne Mahl (German for "beautiful meal"), while dyspepia (Greek for "difficult digestion") refers to its undigested nature. [2] [8]

The well-preserved nature of the specimen suggests that it was eaten by the Compsognathus shortly before the latter's own death and preservation. The disarticulated nature of the specimen suggests that as with some modern predatory birds, Compsognathus may have restrained and dismembered Schoenesmahl during consumption, possibly using its hands and teeth. [2]

Related Research Articles

<i>Archaeopteryx</i> Extinct genus of bird-like dinosaurs

Archaeopteryx, sometimes referred to by its German name, "Urvogel" is a genus of bird-like dinosaurs. The name derives from the ancient Greek ἀρχαῖος (archaīos), meaning "ancient", and πτέρυξ (ptéryx), meaning "feather" or "wing". Between the late 19th century and the early 21st century, Archaeopteryx was generally accepted by palaeontologists and popular reference books as the oldest known bird. Older potential avialans have since been identified, including Anchiornis, Xiaotingia, and Aurornis.

<i>Compsognathus</i> Genus of dinosaurs

Compsognathus is a genus of small, bipedal, carnivorous theropod dinosaur. Members of its single species Compsognathus longipes could grow to around the size of a chicken. They lived about 150 million years ago, during the Tithonian age of the late Jurassic period, in what is now Europe. Paleontologists have found two well-preserved fossils, one in Germany in the 1850s and the second in France more than a century later. Today, C. longipes is the only recognized species, although the larger specimen discovered in France in the 1970s was once thought to belong to a separate species and named C. corallestris.

<i>Struthiomimus</i> Extinct genus of reptile

Struthiomimus, meaning "ostrich-mimic", is a genus of ornithomimid dinosaurs from the late Cretaceous of North America. Ornithomimids were long-legged, bipedal, ostrich-like dinosaurs with toothless beaks. The type species, Struthiomimus altus, is one of the more common, smaller dinosaurs found in Dinosaur Provincial Park; their overall abundance—in addition to their toothless beak—suggests that these animals were mainly herbivorous or omnivorous, rather than purely carnivorous. Similar to the modern extant ostriches, emus, and rheas, ornithomimid dinosaurs likely lived as opportunistic omnivores, supplementing a largely plant-based diet with a variety of small mammals, reptiles, amphibians, insects, invertebrates, and anything else they could fit into their mouth, as they foraged.

<i>Bambiraptor</i> Extinct genus of dinosaurs

Bambiraptor is a Late Cretaceous, 72-million-year-old, bird-like dromaeosaurid theropod dinosaur described by scientists at the University of Kansas, Yale University, and the University of New Orleans.

<i>Procompsognathus</i> Extinct genus of dinosaurs

Procompsognathus is an extinct genus of coelophysid theropod dinosaur that lived approximately 210 million years ago during the later part of the Triassic Period, in what is now Germany. Procompsognathus was a small-sized, lightly built, ground-dwelling, bipedal carnivore, that could grow up to 1 m (3.3 ft) long.

<i>Eustreptospondylus</i> Extinct genus of dinosaurs

Eustreptospondylus is a genus of megalosaurid theropod dinosaur, from the Oxfordian stage of the Late Jurassic period in southern England, at a time when Europe was a series of scattered islands.

<i>Elaphrosaurus</i> Ceratosaurian theropod dinosaur genus from the Late Jurassic Period

Elaphrosaurus is a genus of ceratosaurian theropod dinosaur that lived approximately 154 to 150 million years ago during the Late Jurassic Period in what is now Tanzania in Africa. Elaphrosaurus was a medium-sized but lightly built member of the group that could grow up to 6.2 m (20 ft) long. Morphologically, this dinosaur is significant in two ways. Firstly, it has a relatively long body but is very shallow-chested for a theropod of its size. Secondly, it has very short hindlimbs in comparison with its body. Phylogenetic analyses indicate that this genus is likely a ceratosaur. Earlier suggestions that it is a late surviving coelophysoid have been examined but generally dismissed. Elaphrosaurus is currently believed to be a very close relative of Limusaurus, an unusual beaked ceratosaurian which may have been either herbivorous or omnivorous.

<i>Anoplosaurus</i> Extinct genus of dinosaurs

Anoplosaurus is an extinct genus of herbivorous nodosaurid dinosaur, from the late Albian-age Lower Cretaceous Cambridge Greensand of Cambridgeshire, England. It has in the past been classified with either the armored dinosaurs or the ornithopods, but current thought has been in agreement with the "armored dinosaur" interpretation, placing it in the Ankylosauria.

<i>Sarcolestes</i> Genus of reptiles (fossil)

Sarcolestes is an extinct genus of ankylosaurian ornithischian dinosaur from the Oxford Clay of England. The current type and only species is S. leedsi, and the holotype is a single partial left mandible. The genus and species were named in 1893 by Richard Lydekker, who thought they belonged to a theropod.

<i>Microvenator</i> Extinct genus of dinosaurs

Microvenator is a genus of dinosaur from the Early Cretaceous Cloverly Formation in what is now south central Montana. Microvenator was an oviraptorosaurian theropod. The holotype fossil is an incomplete skeleton, most likely a juvenile with a length of 1.3 m (4.3 ft), and consequently, the adult size remains uncertain. Microvenator celer is primitive and may be the "sister taxon to all other oviraptorosaurs."

<i>Juravenator</i> Extinct genus of dinosaurs

Juravenator is a genus of small coelurosaurian theropod dinosaur, which lived in the area which would someday become the top of the Franconian Jura of Germany, about 151 or 152 million years ago. It is known from a single, juvenile specimen.

Herbstosaurus is the name given to a genus of pterosaurs that lived during the Late Jurassic period, in what is now Argentina. In 1969 Argentine paleobotanist Rafael Herbst in the province Neuquén at Picun Leufú dug up a piece of sandstone holding a number of disarticulated bones of a small reptile. At the time it was assumed the rock dated to the Middle Jurassic (Callovian), about 163 million years ago.

Bavarisaurus is an extinct genus of basal squamate found in the Altmühltal Formation near Bavaria, Germany. It is the only genus in the family Bavarisauridae. A fossil skeleton found in the stomach region of a Compsognathus, a small theropod dinosaur, was originally assigned to the genus by Ostrom (1978), but was renamed to Schoenesmahl by Conrad (2018).

<span class="mw-page-title-main">Compsognathidae</span> Extinct family of dinosaurs

Compsognathidae is a family of coelurosaurian theropod dinosaurs. Compsognathids were small carnivores, generally conservative in form, hailing from the Jurassic and Cretaceous Periods. The bird-like features of these species, along with other dinosaurs such as Archaeopteryx inspired the idea for the connection between dinosaur reptiles and modern-day avian species. Compsognathid fossils preserve diverse integument — skin impressions are known from four genera commonly placed in the group, Compsognathus, Sinosauropteryx, Sinocalliopteryx, and Juravenator. While the latter three show evidence of a covering of some of the earliest primitive feathers over much of the body, Juravenator and Compsognathus also show evidence of scales on the tail or hind legs. "Ubirajara jubatus", informally described in 2020, had elaborate integumentary structures on its back and shoulders superficially similar to the display feathers of a standardwing bird-of-paradise, and unlike any other non-avian dinosaur currently described.

<span class="mw-page-title-main">Opisthotonic death pose</span> Characteristic posture of dinosaur and bird fossils

Non-avian dinosaur and bird fossils are frequently found in a characteristic posture consisting of head thrown back, tail extended, and mouth wide open. The cause of this posture—often called a "death pose"—has been a matter of scientific debate. Traditional explanations ranged from strong ligaments in the animal's neck desiccating and contracting to draw the body into the pose, to water currents arranging the remains in the position.

<i>Sciurumimus</i> Extinct species of reptile

Sciurumimus is an extinct genus of tetanuran theropod from the Late Jurassic Torleite Formation of Germany. It is known from a single juvenile specimen representing the type species, Sciurumimus albersdoerferi, which was found in a limestone quarry close to Painten in Lower Bavaria. The specimen was preserved with traces of feather-like filaments.

The Painten Formation is a geologic formation in Germany. It preserves fossils dating back to the Tithonian stage of the Late Jurassic period.

Bharatagama is an extinct genus of lepidosaur from the Early Jurassic of India. It has been suggested to be one of the oldest known lizards and the oldest known iguanian. The type and only species is Bharatagama rebbanensis, named in 2002. Over one hundred fossils of Bharatagama have been found in the Kota Formation, which outcrops in the Pranhita–Godavari Basin and dates back to about 190 million years ago (Ma). Despite its abundance, Bharatagama is known only from isolated jaw bones mixed together in microvertebrate assemblages with equally fragmentary remains of fish, sphenodontians, dinosaurs, crocodylomorphs, and mammals. These fossils represent all stages of development, from hatchlings to adults. The total length of the skull in adult specimens is estimated to have been about 15 millimetres (0.59 in). Later analysis suggested that the taxon might be a member of Rhynchocephalia.

<span class="mw-page-title-main">Ardeosauridae</span> Extinct family of lizards

Ardeosauridae is an extinct family of lizards known from the Late Jurassic of Germany and North America and Early Cretaceous of Mongolia, with other potential species elsewhere from Europe and Asia over the same time period.

References

  1. Reisdorf, Achim G.; Wuttke, Michael (2012). "Re-evaluating Moodie's Opisthotonic-Posture Hypothesis in Fossil Vertebrates Part I: Reptiles—the taphonomy of the bipedal dinosaurs Compsognathus longipes and Juravenator starki from the Solnhofen Archipelago (Jurassic, Germany)". Palaeobiodiversity and Palaeoenvironments. 92 (1): 119–168. Bibcode:2012PdPe...92..119R. doi:10.1007/s12549-011-0068-y. ISSN   1867-1594.
  2. 1 2 3 4 Conrad, Jack L (2017-12-18). "A new lizard (Squamata) was the last meal of Compsognathus (Theropoda: Dinosauria) and is a holotype in a holotype". Zoological Journal of the Linnean Society. 183 (3): 584–634. doi:10.1093/zoolinnean/zlx055. ISSN   0024-4082.
  3. Wagner, J. A. (1859). "Über einige im lithographischen Schiefer neu aufgefundene Schildkröten und Saurier". Gelehrte Anzeigen der Bayerischen Akademie der Wissenschaften. 49: 553.
  4. Göhlich, Ursula B.; Tischlinger, Helmut; Chiappe, Luis M. (2006). "Juravenator starki (reptilia, theropoda), ein neuer Raubdinosaurier aus dem Oberjura der Südlichen Frankenalb (Süddeutschland)". Archaeopteryx: Jahreszeitschrift der Freunde des Jura-Museums in Eichstätt. 24: 1–26.
  5. Nopcsa, Baron F. (1903). "Neues ueber Compsognathus". Neues Jahrbuch für Mineralogie, Geologie und Paläontologie. 16: 476–494.
  6. Ostrom, J.H. (1978). "The osteology of Compsognathus longipes". Zitteliana. 4: 73–118.
  7. Hoffstetter, R. (1953). Les sauriens anté-cretacées. - Bull. Mus. nat. Hist. natur., 25:345-352, 1 Abb.
  8. Bolet, Arnau; Stubbs, Thomas L; Herrera-Flores, Jorge A; Benton, Michael J (2022-05-03). Zhu, Min; Perry, George H; Zhu, Min (eds.). "The Jurassic rise of squamates as supported by lepidosaur disparity and evolutionary rates". eLife. 11: e66511. doi: 10.7554/eLife.66511 . ISSN   2050-084X. PMC   9064307 . PMID   35502582.