Spongiophyton Temporal range: | |
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Scientific classification ![]() | |
Kingdom: | Plantae |
Division: | incertae sedis |
Family: | † Spongiophytaceae |
Genus: | † Spongiophyton Kräusel (1954) |
Type species | |
†Spongiophyton lenticulare [2] [3] (Barbosa) Kräusel (1954) | |
Species | |
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Spongiophyton was a thallose fossil of the early to mid-Devonian, which is notoriously difficult to classify.
Spongiophyton displayed dichotomous branching, and a flattened/elliptical cross section with a thick (20–80 μm) upper cuticular surface. [4] It is also perforated with pores resembling those of some liverworts. [4] It probably grew on the banks of rivers. [5] Spongiophyton has been mistakenly interpreted as tree resin [6] and lycopod cuticle, [7] and was later identified as the cuticle of a thalloid plant. [8] It has most recently been interpreted on morphological [9] and isotopic [10] grounds as a lichen—which would place it with Winfrenatia among the earliest known representatives of this group. [11]
The significance of the isotopic data has, however, been called into question. Jahren et al. argued that mosses and liverworts had a δ13C signature of under −26‰, and lichens were exclusively > −26‰. But in deducing this they relied solely on their own data, neglecting to include published datasets or bryophytes from a wide range of habitats. They also failed to take into account any adjustment necessary to overcome post-burial alteration of the δ13C, or to compensate for the different isotopic composition of the early Devonian atmosphere. [4] Repeating Jahren's experiments with these factors taken into account shows that most major groups' δ13C values overlap significantly, and do not provide a statistically significant case for the inclusion of Spongiophyton in any group. [4]
The Ediacaran is a geological period of the Neoproterozoic era that spans 96 million years from the end of the Cryogenian period at 635 Mya to the beginning of the Cambrian period at 538.8 Mya. It is the last period of the Proterozoic eon as well as the last of the so-called "Precambrian supereon", before the beginning of the subsequent Cambrian period marks the start of the Phanerozoic eon, where recognizable fossil evidence of life becomes common.
The cloudinids, an early metazoan family containing the genera Acuticocloudina, Cloudina and Conotubus, lived in the late Ediacaran period about 550 million years ago. and became extinct at the base of the Cambrian. They formed millimetre-scale conical fossils consisting of calcareous cones nested within one another; the appearance of the organism itself remains unknown. The name Cloudina honors the 20th-century geologist and paleontologist Preston Cloud.
A lichen is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship. Lichens are important actors in nutrient cycling and act as producers which many higher trophic feeders feed on, such as reindeer, gastropods, nematodes, mites, and springtails. Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not plants. They may have tiny, leafless branches (fruticose); flat leaf-like structures (foliose); grow crust-like, adhering tightly to a surface (substrate) like a thick coat of paint (crustose); have a powder-like appearance (leprose); or other growth forms.
Rangea is a frond-like Ediacaran fossil with six-fold radial symmetry. It is the type genus of the rangeomorphs.
Cholestane is a saturated tetracyclic triterpene. This 27-carbon biomarker is produced by diagenesis of cholesterol and is one of the most abundant biomarkers in the rock record. Presence of cholestane, its derivatives and related chemical compounds in environmental samples is commonly interpreted as an indicator of animal life and/or traces of O2, as animals are known for exclusively producing cholesterol, and thus has been used to draw evolutionary relationships between ancient organisms of unknown phylogenetic origin and modern metazoan taxa. Cholesterol is made in low abundance by other organisms (e.g., rhodophytes, land plants), but because these other organisms produce a variety of sterols it cannot be used as a conclusive indicator of any one taxon. It is often found in analysis of organic compounds in petroleum.
The Ediacaranbiota is a taxonomic period classification that consists of all life forms that were present on Earth during the Ediacaran Period. These were enigmatic tubular and frond-shaped, mostly sessile, organisms. Trace fossils of these organisms have been found worldwide, and represent the earliest known complex multicellular organisms. The term "Ediacara biota" has received criticism from some scientists due to its alleged inconsistency, arbitrary exclusion of certain fossils, and inability to be precisely defined.
Prototaxites is an extinct genus of terrestrial fossil fungi dating from the Late Silurian until the Late Devonian periods. Prototaxites formed large trunk-like structures up to 1 metre (3 ft) wide, reaching 8 metres (26 ft) in length, made up of interwoven tubes around 50 micrometres (0.0020 in) in diameter, making it by far the largest land-dwelling organism of its time.
Nematothallus is a form genus comprising cuticle-like fossils. Some of its constituents likely represent red algae, whereas others resemble lichens.
Cosmochlaina is a form genus of nematophyte – an early plant known only from fossil cuticles, often found in association with tubular structures. The form genus was put forwards by Dianne Edwards, and is diagnosed by inwards-pointing flanges and randomly oriented pseudo-cellular units. Projections on the outer surface are always present, and sometimes also appear on the inner surface; however, the surface of the cuticle itself is always smooth. The holes in the cuticle are often covered by round flaps, loosely attached along a side.
Euthycarcinoidea are an enigmatic group of extinct possibly amphibious arthropods that ranged from Cambrian to Triassic times. Fossils are known from Europe, North America, Argentina, Australia and Antarctica.
Ernietta is an extinct genus of Ediacaran organisms with an infaunal lifestyle. Fossil preservations and modeling indicate this organism was sessile and “sack”-shaped. It survived partly buried in substrate, with an upturned bell-shaped frill exposed above the sediment-water interface. Ernietta have been recovered from present-day Namibia, and are a part of the Ediacaran biota, a late Proterozoic radiation of multicellular organisms. They are among the earliest complex multicellular organisms and are known from the late Ediacaran. Ernietta plateauensis remains the sole species of the genus.
In geochemistry, paleoclimatology, and paleoceanography δ13C is an isotopic signature, a measure of the ratio of the two stable isotopes of carbon—13C and 12C—reported in parts per thousand. The measure is also widely used in archaeology for the reconstruction of past diets, particularly to see if marine foods or certain types of plants were consumed.
The end-Ediacaran extinction is a mass extinction believed to have occurred near the end of the Ediacaran period, the final period of the Proterozoic eon. Evidence suggesting that such a mass extinction occurred includes a massive reduction in diversity of acritarchs, the sudden disappearance of the Ediacara biota and calcifying organisms, and the time gap before Cambrian organisms "replaced" them. Some lines of evidence suggests that there may have been two distinct pulses of the extinction event, one occurring 550 million years ago and the other 539 million years ago.
The end-Botomian mass extinction event, also known as the late early Cambrian extinctions, refer to two extinction intervals that occurred during Stages 4 and 5 of the Cambrian Period, approximately 513 to 509 million years ago. Estimates for the decline in global diversity over these events range from 50% of marine genera up to 80%. Among the organisms affected by this event were the small shelly fossils, archaeocyathids, trilobites, brachiopods, hyoliths, and mollusks.
Gregory John Retallack is an Australian paleontologist, geologist, and author who specializes in the study of fossil soils (paleopedology). His research has examined the fossil record of soils though major events in Earth history, extending back some 4.6 billion years. Among his publications he has written two standard paleopedology textbooks, said N. Jones in Nature Geoscience "Retallack has literally written the book on ancient soils."
Protonympha is a form genus for problematic fossils of Devonian age in New York. It has been of special interest because of its morphological similarity with the iconic Ediacaran fossil Spriggina, and may have been a late surviving vendobiont.
Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils. This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2019.
The Shuram excursion, or Shuram-Wonoka excursion, is a change in δ13C, or in the ratio of carbon-13 to carbon-12, taking place between around 573 and 562 million years ago, during the Ediacaran Period. It was first noticed in the Wonoka Formation in South Australia in 1990 and later in the Shuram Formation in Oman in 1993. It is the largest negative δ13C excursion in Earth history, and recovery took 50 million years, although the apparent magnitude of the excursion may be distorted due to meteoric water diagenesis.
Cuticle analysis, also known as fossil cuticle analysis and cuticular analysis, is an archaeobotanical method that uses plant cuticles to reconstruct the vegetation of past grassy environments. Cuticles comprise the protective layer of the skin, or epidermis, of leaves and blades of grass. They are made of cutin, a resilient substance that can preserve the shapes of underlying cells, a quality that aids in the identification of plants that are otherwise no longer visible in the archaeological record. This can inform archaeobotanists on the floral makeup of a past environment, even when surviving remains from the plants are limited. Plant cuticles have also been incorporated into other areas of archaeobotanical research based on their susceptibility to environmental factors such as pCO2 levels and stresses such as water deficit and sodium chloride exposure. Such research can help to reconstruct past environments and identify ecological events.
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