Tannuella Temporal range: | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Mollusca |
Class: | † Helcionelloida |
Order: | † Helcionelliformes |
Family: | † Helcionellidae |
Genus: | † Tannuella |
Tannuella is a genus of helcionellids known from Lower Cambrian strata.
It has a high, septate shell similar to that of Knightoconus . These genera were once considered ancestral cephalopods, [1] but are more broadly assigned to the helcionellids (or putatively the tergomyans); i.e. "Cambrian monoplacophora". [2]
Nectocaris is a genus of squid-like animal of controversial affinities known from the Cambrian period. The initial fossils were described from the Burgess Shale of Canada. Other similar remains possibly referrable to the genus are known from the Emu Bay Shale of Australia and Chengjiang Biota of China.
Coleoidea or Dibranchiata is one of the two subclasses of cephalopods containing all the various taxa popularly thought of as "soft-bodied" or "shell-less". Unlike its extant sister group Nautiloidea, whose members have a rigid outer shell for protection, the coleoids have at most an internal shell called cuttlebone or gladius that is used for buoyancy or as muscle anchorage. Some species, notably incirrate octopuses, have lost their cuttlebone altogether, while in some it has been replaced by a chitinous support structure. A unique trait of the group is the ability to edit their own RNA.
Nautiloids are a group of marine cephalopods (Mollusca) which originated in the Late Cambrian and are represented today by the living Nautilus and Allonautilus. Fossil nautiloids are diverse and species rich, with over 2,500 recorded species. They flourished during the early Paleozoic era, when they constituted the main predatory animals. Early in their evolution, nautiloids developed an extraordinary diversity of shell shapes, including coiled morphologies and giant straight-shelled forms (orthocones). No orthoconic and only a handful of coiled species, the nautiluses, survive to the present day.
The evolution of the molluscs is the way in which the Mollusca, one of the largest groups of invertebrate animals, evolved. This phylum includes gastropods, bivalves, scaphopods, cephalopods, and several other groups. The fossil record of mollusks is relatively complete, and they are well represented in most fossil-bearing marine strata. Very early organisms which have dubiously been compared to molluscs include Kimberella and Odontogriphus.
Latouchella is an extinct genus of marine invertebrate animal, that is considered to be a mollusk and which may be a sea snail, a gastropod. It is a helcionellid from the Tommotian epoch of what is now Siberia. Its tightly coiled, spiral shell contains a number of low "walls" running up the front surface of the interior; these would have directed water currents within its shell. Between these walls are a series of furrows, parallel to the shell's aperture, giving casts of the internal structure the appearance of a railway line, with sleepers tying together paired rails that run towards the apex of the shell.
Plectronocerida is a primitive order from which subsequent cephalopod orders are ultimately derived.
The Ellesmerocerida is an order of primitive cephalopods belonging to the subclass Nautiloidea with a widespread distribution that lived during the Late Cambrian and Ordovician.
Scenella is an extinct genus of fossil invertebrate animal which is generally considered to be a mollusc; at various times it has been suggested that this genus belongs with the gastropods, the monoplacophorans, or the helcionellids, although no firm association with any of these classes has been established. An affinity with the hydrozoa has been considered, although some authors oppose this hypothesis. A gastropod affinity is defended on the basis of six pairs of internal muscle scars, whilst the serially-repeated nature of these scars suggests to other authors a monoplacophoran affinity. However the specimens showing this scarring have not been convincingly shown to belong to the genus Scenella. A similarity to the Ediacaran Ovatoscutum has also been drawn.
Helcionellid or Helcionelliformes is an order of small fossil shells that are universally interpreted as molluscs, though no sources spell out why this taxonomic interpretation is preferred. These animals are first found about 540 to 530 million years ago in the late Nemakit-Daldynian age, which is the earliest part of the Cambrian period. A single species persisted to the Early Ordovician. These fossils are component of the small shelly fossils (SSF) assemblages.
Mollusca is the second-largest phylum of invertebrate animals, after Arthropoda; members are known as molluscs or mollusks. Around 76,000 extant species of molluscs are recognized. The number of fossil species is estimated between 60,000 and 100,000 additional species. The proportion of undescribed species is very high. Many taxa remain poorly studied.
Plectronoceras is the earliest known shelled cephalopod, dating to the Late Cambrian. None of the fossils are complete, and none show the apex or aperture of the shell. Approximately half of its shell was filled with septa; 7 were recorded in a 2 centimetres (0.79 in) shell. Its shell contains transverse septa separated by about half a millimetre, with a siphuncle on its concave side. Its morphology matches closely to that hypothesised for the last common ancestor of all cephalopods.
Knightoconus antarcticus is an extinct species of fossil monoplacophoran from the Cambrian Minaret Formation of Antarctica. It is thought to represent an ancestor to the cephalopods. It had a chambered conical shell, but lacked a siphuncle.
Volborthella is an animal of uncertain classification, whose fossils pre-date 530 million years ago. It has been considered for a period a cephalopod. However discoveries of more detailed fossils showed that Volborthella’s small, conical shell was not secreted but built from grains of the mineral silicon dioxide (silica), and that it was not divided into a series of compartments by septa as those of fossil shelled cephalopods and the living Nautilus are. This illusion was a result of the laminated texture of the organisms' tests. Therefore, Volborthella’s classification is now uncertain. It has been speculated that it may in fact represent a sclerite of a larger organism, on the basis of one specimen; however, it may be premature to accept this hypothesis, as the arrangement of sclerites producing this impression may have occurred by chance. The Ordovician scleritome-bearing Curviconophorus, as well as the Halwaxiids, lobopods and echinoderms, demonstrate the diversity of organisms which may produce a scleritome of this nature. The related Campitius was originally suggested to be part of a radula rather than a scleritome, but is now considered a synonym of Volborthella.
The cephalopods have a long geological history, with the first nautiloids found in late Cambrian strata, and purported stem-group representatives present in the earliest Cambrian lagerstätten.
Plectronoceratoidea is a superorder or subclass containing primitive nautiloids from the Late Cambrian and Early Ordovician. This group is best considered a paraphyletic grade of early cephalopods, as it contains the ancestors of subsequent post-Cambrian cephalopod orders.
Shelbyoceras is a genus of Cambrian monoplacophora which was one of the genera mistaken for a cephalopod, since the characteristics differentiating monoplacophora from cephalopods are few. Shelbyoceras was reclassified based on a depressed groove that forms a band around the shell, which is similar to a feature seen in Hypseloconus. The septa in this genus are either closely or irregularly spaced.
Eotebenna is a genus of helcionellid from the Early to Middle Cambrian of Greenland, Denmark and Australia, with sail-like "snorkels", similar to Latouchella.
Eurekapegma is a helcionellid from the Middle Cambrian of New Zealand. It is flattened sideways, with a dividing wall within its shell; it resembles the helcionellid Eotebenna.
Protowenella is a genus of helcionellid from the Middle Cambrian of Australia. It has a strongly spiralled, smooth shell with concentric ridges that have low relief. Other than its tighter coiling, it is very similar to Tichkaella.
Helcionella is a genus of helcionellid, a fossil marine invertebrate animal that is considered to be a mollusk and may possibly be a gastropod. The shells of these animals are about a centimetre in size. The tip of the shell extends beyond the rear extremity of the shell's aperture, and the shell is "endogastric" in shape.