Tisamenus clotho

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Tisamenus clotho
Tisamenus cf. clotho 'Camarines'.JPG
Tisamenus clotho 'Camarines Norte', pair
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Phasmatodea
Family: Heteropterygidae
Subfamily: Obriminae
Tribe: Obrimini
Genus: Tisamenus
Species:
T. clotho
Binomial name
Tisamenus clotho
Synonyms [1]
  • Hoploclonia clothoRehn, J.A.G. & Rehn, J.W.H., 1939

Tisamenus clotho is a stick insect species native to the Philippines.

Contents

Eggs in dorsal and lateral view Tisamenus clotho eggs.jpg
Eggs in dorsal and lateral view

Description

Females of Tisamenus clotho are light brown to buckthorn brown and often show other shades of brown, with the head and legs being darker. They reach lengths about 56 millimetres (2.2 in). Males are significantly smaller and dark brown. The head is almost square and only slightly longer than wide. The supraorbitals are formed as three prominent, elongated tubercles. The occipitals and median coronals are present as flat tubercles, the lateral coronals are bifid, rounded tubercles. As with Tisamenus deplanatus, the triangular region typical of the genus on the mesonotum is relatively short, reaching just under the middle of the mesothorax, where it forms an approximately equilateral triangle forms. From the center of the base of the triangle at the anterior edge of the mesonotum, a faintly indicated carina runs across the triangle to its posterior angle. From there it extends as a prominent longitudinal carina further over the rest of the mesonotum and the entire metanotum. The mesopleura are armed with four lateral spines, the metapleura with two lateral spines. Behind are the supracoxals, which are also spiny and whose supracoxal angle is shaped into a short tubercle. [2]

Distribution

The type material of the species comes from Polillo Island. [2] The animals being bred were collected in two localities in the province of Camarines Norte in southern Luzon. These are located at Mount Bagacay in the Barangay Fundado of Labo and at Mananap falls in the Barangay Fabrica of Daet. [1] [3]

Taxonomy

James Abram Garfield Rehn and his son John William Holman Rehn described the species in 1939 under the basionym Hoploclonia clotho. An adult female collected by Taylor on Polillo Island from the collection of Morgan Hebard at the Academy of Natural Sciences of Drexel University in Philadelphia was chosen as holotype. From the collection of the United States National Museum, a juvenile male has been chosen as allotype and an adult female as another paratype. In addition, Rehn and Rehn examined three other juvenile females from this collection and one juvenile male from the Hebard collection for their description. All specimens examined were from Polillo. They divided the Philippine species of the genus into different groups according to morphological features. Hoploclonia clotho was placed with Hoploclonia serratoria (today Tisamenus serratorius ), Hoploclonia asper (today Tisamenus asper ), Hoploclonia atropis (today Tisamenus atropis ) in the so-called Serratoria group. Their species are relatively spiny, have distinct lateral spines along the margins of the meso- and metathorax and an isosceles triangle on the anterior mesothorax, reaching about halfway down the mesonotum. [2] Since Oliver Zompro transferred the Philippine species of Hoploclonia to the genus Tisamenus in 2004 and left only those occurring on Borneo in the genus Hoploclonia, the species is named Tisamenus clotho. [1] [4]

In a molecular genetics study published in 2021, representatives of other Tisamenus species were included in addition to Tisamenus clotho from Camarines Norte. It turned out that the species is not so closely related to Tisamenus serratorius, as Rehn and Rehn had suspected in their 1939 group classification. It is more closely related to Tisamenus deplanatus and two undescribed or unidentified species from Camiguin and Sibuyan. [3]

In captivity

A stock keeping by enthusiasts in terrariums derives from specimens from Camarines Norte collected by Thierry Heitzmann on July 22 at Mount Bagacay and August 4, 2015 at Mananap falls. Their species affiliation was not certain at first, so they were named Tisamenus cf. clotho 'Camarines'. Joachim Bresseel identified these as Tisamenus clotho. The species is easy to keep and breed. They eat leaves of bramble, other Rosaceae and hazel. [3] [5]

Related Research Articles

<span class="mw-page-title-main">Heteropterygidae</span> Family of stick insects

The Heteropterygidae is a family of stick insects belonging to the suborder Euphasmatodea. Species can be found in Australasia, East and Southeast Asia. About 150 valid species have been described.

<span class="mw-page-title-main">Obriminae</span> Family of stick insects

The Obriminae are the most species-rich subfamily of the Phasmatodea family Heteropterygidae native to Southeast Asia. It is divided into two tribe.

<i>Hoploclonia</i> Tribe of stick insects

Hoploclonia is the only genus of the tribe Hoplocloniini and brings together relatively small and darkly coloured Phasmatodea species.

<i>Tisamenus</i> (insect) Genus of stick insects

The genus Tisamenus native to the Philippines combines small to medium-sized species of stick insects.

<i>Eubulides</i> (insect) Genus of stick insects

Eubulides is a stick insect genus native to the Philippines.

<i>Tisamenus serratorius</i> Species of stick insect

Tisamenus serratorius is a stick insect species that occurs on the Philippine island Luzon.

<i>Tisamenus deplanatus</i> Species of stick insect

Tisamenus deplanatus is a stick insect species native to the Philippine islands Luzon and Mindanao occurs.

<i>Tisamenus fratercula</i> Species of stick insect

Tisamenus fratercula is a stick insect species native to the Philippine island Luzon.

<i>Heterocopus</i> Genus of stick insects

Heterocopus is a monotypic stick insect genus, containing Heterocopus leprosus as the only valid species.

<i>Stenobrimus</i> Genus of stick insects

Stenobrimus is a genus of medium-sized stick insects native to the Philippines.

Pterobrimus is a monotypic genus of stick insects (Phasmatodea), containing the species Pterobrimus depressus, which is native to Fiji.

<i>Tisamenus draconinus</i> Species of stick insect

Tisamenus draconinus is a species of stick insect in the family Heteropterygidae native to the Philippines.

<i>Tisamenus ranarius</i> Species of stick insect

Tisamenus ranarius is a stick insect species (Phasmatodea), in the family of the Heteropterygidae native to the Philippines.

<i>Tisamenus hebardi</i> Species of stick insect

Tisamenus hebardi is a stick insect species (Phasmatodea), in the family of the Heteropterygidae endemic to the north of the Philippine island of Luzon.

<i>Tisamenus armadillo</i> Species of stick insect

Tisamenus armadillo is a stick insect species (Phasmatodea) in the family Heteropterygidae, endemic to the Philippines.

<i>Tisamenus hystrix</i> Species of stick insect

Tisamenus hystrix is a stick insect species (Phasmatodea), in the family of the Heteropterygidae endemic to the Philippines.

<i>Tisamenus lachesis</i> Species of stick insect

Tisamenus lachesis is a stick insect species (Phasmatodea), in the family of the Heteropterygidae endemic to the Philippine Polillo Island.

Tisamenus cervicornis is a stick insect species (Phasmatodea), in the family of the Heteropterygidae endemic to the Philippine island Luzon in the province of Camarines Sur.

Tisamenus asper is a stick insect species (Phasmatodea), in the family of the Heteropterygidae endemic to the Philippine island Luzon.

<i>Eubulides igorrote</i> Species of stick insect

Eubulides igorrote is a stick insect species from the family of the Heteropterygidae native to the Philippine island of Luzon.

References

  1. 1 2 3 Brock, P. D.; Büscher, T. H. & Baker, E. W.: Phasmida Species File Online . Version 5.0./5.0. (accessdate 5 August 2022)
  2. 1 2 3 Rehn, J. A. G. & Rehn, J. W. H. (1939). Proceedings of The Academy of Natural Sciences (Vol. 90, 1938) , Philadelphia, pp. 466–468 & pp. 473-475; pl. 34 fig. 29
  3. 1 2 3 Bank, S.; Buckley, T. R.; Büscher, T. H.; Bresseel, J.; Constant, J.; de Haan, M.; Dittmar, D.; Dräger, H.; Kahar, R. S.; Kang, A.; Kneubühler, B.; Langton-Myers, S. & Bradler, S. (2021). Reconstructing the nonadaptive radiation of an ancient lineage of ground-dwelling stick insects (Phasmatodea: Heteropterygidae) , Systematic Entomology, DOI: 10.1111/syen.12472
  4. Zompro, O. (2004). Revision of the genera of the Areolatae, including the status of Timema and Agathemera (Insecta, Phasmatodea), Goecke & Evers, Keltern-Weiler, pp. 200–207, ISBN   978-3931374396
  5. Information about Tisamenus cf. clotho 'Camarines' on Phasmatodea.com by Hennemann, F. H.; Conle, O. V.; Kneubühler, B. & Valero, P.