Townsend's ground squirrel | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Rodentia |
Family: | Sciuridae |
Genus: | Urocitellus |
Species: | U. townsendii |
Binomial name | |
Urocitellus townsendii (Bachman, 1839) | |
Synonyms | |
Spermophilus townsendiiBachman, 1839 |
Townsend's ground squirrel (Urocitellus townsendii) is a species of rodent in the family Sciuridae. It is found in high desert shrublands in several areas of the United States.
Townsend's ground squirrel is found in the Great Basin and the Columbia Plateau. Its range includes south-central Washington, eastern Oregon, southern Idaho, western Utah, most of Nevada, and extreme eastern California. [2] The subspecies are distributed as follows: [3]
Townsend's ground squirrels typically inhabit arid grasslands and shrub-grasslands. Malheur ground squirrels, however, inhabit the relatively mesic and fertile Snake River Plain. [2] Plant communities in which Townsend's ground squirrels occur include crested wheatgrass ( Agropyron cristatum ), big sagebrush ( Artemisia tridentata ) wheatgrass, [4] rabbitbrush ( Chrysothamnus spp.), shadscale (Atriplex confertifolia), and winterfat (Kraschenninikovia lanata) grasslands. [5]
In southeastern Idaho, the density of Townsend's ground squirrel burrows is highest on winterfat-Sandberg bluegrass ( Poa secunda ), intermediate in big sagebrush-Thurber needlegrass ( Stipa thurberiana ), and lowest in shadscale-Indian ricegrass ( Oryzopsis hymenoides ) and thread grass ( Stipa comata ) . Burrow densities were highly variable in mixed exotic annual communities, and negatively correlated with cheatgrass (Bromus tectorum) . They increase with increasing native grass cover, but the populations are unstable when native grass cover is overly high. [6] [7]
Although Townsend's ground squirrels occur in arid environments, within those environments they are most common around desert springs and irrigated fields. [8] They also occupy ridgetops, hillsides, and valley bottoms, [2] canal and railroad embankments, and old fields. [9] As a burrowing species, Townsend's ground squirrels select sites with deep, friable, well-drained soils. [2] [5] In southeastern Idaho, 68% of Townsend's ground squirrel burrows were in sand, 28% in silt, and 4% in clay. [10]
Smith and Johnson reported a mean home range of 1,357 square meters for 14 Snake Valley ground squirrels. [11] Townsend's ground squirrel density can fluctuate greatly from year to year. Estimated population density of Snake Valley ground squirrels ranged from 3 to 32 individuals per hectare. [11] Densities of 296 to 331 individuals per hectare have been reported for Piute ground squirrels. [12]
Townsend's ground squirrels become dormant in late spring or early summer, after grasses cure. They emerge from dormancy in late winter. Dormancy lasts 7.5 to 9 continuous months. [12] [13] It is shorter in wet years, when green forage is available later in summer, than in dry years. [11] [12]
Females breed as yearlings. Most males also breed as yearlings, although male Snake Valley ground squirrels breed at 2 years of age. [11] [13] Most breeding occurs in late January or early February, [11] just after dormancy ends. Piute ground squirrels breed from mid-February to early March, somewhat later than other subspecies. [13] [14] Gestation is about 24 days. [12] One litter is produced per year, with 6 to 10 pups per litter. [2] Pups are born hairless and with eyes closed. Early postnatal development of Townsend's ground squirrels is relatively slow compared to development of other Spermophilus species. Pups open their eyes at 19 to 22 days of age, and are weaned shortly thereafter. [2]
Townsend's ground squirrels occupy open habitats and use burrows for shelter, protection from predators, and food storage. Burrows are often grouped into colonies, but some Townsend's ground squirrels are solitary. [2] Except when mothers have pups, there is only one Townsend's ground squirrel per burrow. Burrows have one to many openings and may have numerous auxiliary burrows in addition to the "home" or nest burrow. [7] Burrow dimensions of Townsend's ground squirrels in southeastern Idaho ranged from 2.6 to 3.8 inches (6.5–9.6 cm) horizontally and from 1.7 to 2.5 inches (4.3–6.3 cm) vertically. [10]
Townsend's ground squirrels have been observed climbing shrubs while foraging, apparently for cover and to spot palatable vegetation. [2]
Townsend's ground squirrels consume mainly green vegetation and some seeds and insects. [15] [16] Green grasses are apparently a staple from late winter until just prior to grass senescence and Townsend ground squirrel estivation, when seeds become the primary diet item. Seeds are an important source of calories just prior to estivation. [14] Where present, winterfat is browsed heavily, [17] but only light browsing of other shrubs has been reported. From March through May on the Arid Land Ecology Reserve in eastern Washington, the Townsend's ground squirrel diet is 49% Sandberg bluegrass, 11% western yarrow (Achillea millefolium var. occidentalis), 8% pinnate tansymustard (Descurania pinnata) seed, 31% other plant species (mostly forbs), and 1% insects. [16] On a big sagebrush-crested wheatgrass community in southeastern Idaho, 80% of Townsend's ground squirrels trapped in June had consumed crested wheatgrass, and Townsend's ground squirrels became dormant after the crested wheatgrass senesced. [18] Cheatgrass (Bromus tectorum) is an important food item in some years. As an annual with wide year-to-year swings in productivity, however, it is not a reliable food source. [7]
Adult cannibalism of unweaned young has been observed in the Townsend's ground squirrel. [19]
Townsend's ground squirrels are the primary prey of ferruginous hawks ( Buteo regalis ) in the Great Basin and Columbia Plateau. [4] They are also a major and often primary diet item of prairie falcons ( Falco mexicanus ). [5] [20] The Townsend's ground squirrel has been rated one of the two most important prey species in southeastern Idaho because of its importance to ferruginous hawks and prairie falcons. [5] Other important predators of Townsend's ground squirrels include other hawks (Accipiter and Buteo spp.) and falcons (Falco spp.), common crows (Corvus corax), badgers (Taxidea taxus), coyotes (Canis latrans), long-tailed weasels (Mestrela frenata), western rattlesnakes (Crotalus viridis), and gopher snakes (Pituophis melanoleucus). [2] [11] [12] [21]
The Great Basin Desert is part of the Great Basin between the Sierra Nevada and the Wasatch Range. The desert is a geographical region that largely overlaps the Great Basin shrub steppe defined by the World Wildlife Fund, and the Central Basin and Range ecoregion defined by the U.S. Environmental Protection Agency and United States Geological Survey. It is a temperate desert with hot, dry summers and snowy winters. The desert spans large portions of Nevada and Utah, and extends into eastern California. The desert is one of the four biologically defined deserts in North America, in addition to the Mojave, Sonoran, and Chihuahuan Deserts.
The northern Idaho ground squirrel is a species of the largest genus of ground squirrels. This species and the Southern Idaho ground squirrel were previously considered conspecific, together called the Idaho ground squirrel.
The pygmy rabbit is a rabbit species native to the United States. It is also the only native rabbit species in North America to dig its own burrow. The pygmy rabbit differs significantly from species within either the Lepus (hare) or Sylvilagus (cottontail) genera and is generally considered to be within the monotypic genus Brachylagus. One isolated population, the Columbia Basin pygmy rabbit, is listed as an endangered species by the U.S. Federal government, though the International Union for Conservation of Nature lists the species as lower risk.
The Washington ground squirrel is near threatened species of squirrel distributed in the Pacific Northwest states of Washington and Oregon, United States.
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Belding's ground squirrel, also called pot gut, sage rat or picket-pin, is a squirrel that lives on mountains in the western United States. In California, it often is found at 6,500 to 11,800 feet (2,000–3,600 m) in meadows between Lake Tahoe and Kings Canyon. This species is not of conservation concern, and its range includes some protected areas.
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The Blue Mountains ecoregion is a Level III ecoregion designated by the United States Environmental Protection Agency (EPA) in the Pacific Northwest, mainly in the state of Oregon, with small areas over the state border in Idaho and southeastern Washington. It is also contiguous with the World Wildlife Fund's Blue Mountain forests ecoregion.
The Snake River Plain ecoregion is a Level III ecoregion designated by the United States Environmental Protection Agency (EPA) in the U.S. states of Idaho and Oregon. It follows the Snake River across Idaho, stretching roughly 400 miles (640 km) from the Wyoming border to Eastern Oregon in the xeric intermontane west. Characterized by plains and low hills, it is considerably lower and less rugged than surrounding ecoregions. Many of the alluvial valleys bordering the Snake River are used for agriculture. Where irrigation water and soil depth are sufficient, sugar beets, potatoes, alfalfa, small grains, and vegetables are grown. Elsewhere, livestock grazing is widespread. Cattle feedlots and dairy operations are found locally.
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Urocitellus is a genus of ground squirrels. They were previously believed to belong to the much larger genus Spermophilus, but DNA sequencing of the cytochrome b gene showed that this group was paraphyletic to the prairie dogs and marmots, and could therefore no longer be retained as a single genus. As a result, Urocitellus is now considered as a genus in its own right.
This article incorporates public domain material from Spermophilus townsendii. United States Department of Agriculture.