Marmots Temporal range: Late Miocene – recent | |
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Alpine marmot (Marmota marmota) | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Rodentia |
Family: | Sciuridae |
Tribe: | Marmotini |
Genus: | Marmota Blumenbach, 1779 |
Type species | |
Marmota marmota | |
Species | |
15, see text |
Marmots are large ground squirrels in the genus Marmota, with 15 species living in Asia, Europe, and North America. These herbivores are active during the summer, when they can often be found in groups, but are not seen during the winter, when they hibernate underground. They are the heaviest members of the squirrel family. [1]
Marmots are large rodents with characteristically short but robust legs, enlarged claws which are well adapted to digging, stout bodies, and large heads and incisors to quickly process a variety of vegetation. While most species are various forms of earthen-hued brown, marmots vary in fur coloration based roughly on their surroundings. Species in more open habitat are more likely to have a paler color, while those sometimes found in well-forested regions tend to be darker. [2] [3] Marmots are the heaviest members of the squirrel family. Total length varies typically from about 42 to 72 cm (17 to 28 in) and body mass averages about 2 kg (4+1⁄2 lb) in spring in the smaller species and 8 kg (18 lb) in autumn, at times exceeding 11 kg (24 lb), in the larger species. [4] [5] [6] The largest and smallest species are not clearly known. [3] [4] In North America, on the basis of mean linear dimensions and body masses through the year, the smallest species appears to be the Alaska marmot and the largest is the Olympic marmot. [5] [7] [8] [6] Some species, such as the Himalayan marmot and Tarbagan marmot in Asia, appear to attain roughly similar body masses to the Olympic marmot, but are not known to reach as high a total length as the Olympic species. [9] [10] In the traditional definition of hibernation, the largest marmots are considered the largest "true hibernators" (since larger "hibernators" such as bears do not have the same physiological characteristics as obligate hibernating animals such as assorted rodents, bats and insectivores). [11] [12]
Some species live in mountainous areas, such as the Alps, northern Apennines, Carpathians, Tatras, and Pyrenees in Europe; northwestern Asia; the Rocky Mountains, Black Hills, the Cascade and Pacific Ranges, and the Sierra Nevada in North America; and the Deosai Plateau in Pakistan and Ladakh in India. Other species prefer rough grassland and can be found widely across North America and the Eurasian Steppe. The slightly smaller and more social prairie dog is not classified in the genus Marmota, but in the related genus Cynomys.
Marmots typically live in burrows (often within rockpiles, particularly in the case of the yellow-bellied marmot), and hibernate there through the winter. Most marmots are highly social and use loud whistles to communicate with one another, especially when alarmed.
Marmots mainly eat greens and many types of grasses, berries, lichens, mosses, roots, and flowers.
The following is a list of all Marmota species recognized by Thorington and Hoffman [13] plus the recently defined M. kastschenkoi. [14] They divide marmots into two subgenera.
Subgenus | Image | Name | Common Name | Distribution |
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Marmota | Marmota baibacina | Gray marmot or Altai marmot | Siberia | |
Marmota bobak | Bobak marmot | eastern Europe to central Asia | ||
Marmota broweri | Alaska marmot, Brower's marmot, or Brooks Range marmot | Alaska | ||
Marmota camtschatica | Black-capped marmot | eastern Siberia | ||
Marmota caudata | Long-tailed marmot, golden marmot, or red marmot | central Asia | ||
Marmota himalayana | Himalayan marmot or Tibetan snow pig | the Himalayas | ||
Marmota kastschenkoi | Forest-steppe marmot | south Russia [14] | ||
Marmota marmota | Alpine marmot | Europe only in the Alps, Carpathian Mountains, Tatra Mountains, northern Apennine Mountains, and reintroduced in the Pyrenees | ||
Marmota menzbieri | Menzbier's marmot | central Asia | ||
Marmota monax | Groundhog, woodchuck, or whistlepig | Canada and east of the Mississippi in northern USA | ||
Marmota sibirica | Tarbagan marmot, Mongolian marmot, or tarvaga | Siberia | ||
Petromarmota | Marmota caligata | Hoary marmot | northwestern North America (Alaska, Yukon, British Columbia, Alberta, Washington, Montana) | |
Marmota flaviventris | Yellow-bellied marmot | southwestern Canada and western United States | ||
Marmota olympus | Olympic marmot | endemic to the Olympic Peninsula, Washington, United States | ||
Marmota vancouverensis | Vancouver Island marmot | endemic to Vancouver Island, British Columbia, Canada | ||
Some extinct species of marmots are recognized from the fossil record, for example:
Marmots have been known since antiquity. Research by the French ethnologist Michel Peissel claimed the story of the "Gold-digging ant" reported by the Ancient Greek historian Herodotus, who lived in the fifth century BCE, was founded on the golden Himalayan marmot of the Deosai Plateau and the habit of local tribes such as the Brokpa to collect the gold dust excavated from their burrows. [19] Some historians believe that Strabo's λέων μύρμηξ and Agatharchides's μυρμηκολέων, most probably are the marmot. [20]
An anatomically accurate image of a marmot was printed and distributed as early as 1605 by Jacopo Ligozzi, who was noted for his images of flora and fauna.
The etymology of the term "marmot" is uncertain. It may have arisen from the Gallo-Romance prefix marm-, meaning to mumble or murmur (an example of onomatopoeia). Another possible origin is postclassical Latin, mus montanus, meaning "mountain mouse". [21]
Beginning in 2010, Alaska celebrates February 2 as "Marmot Day", a holiday intended to observe the prevalence of marmots in that state and take the place of Groundhog Day. [22]
Some historians and paleogeneticists have postulated that the Yersinia pestis variant that caused the Black Death pandemic that struck Eurasia in the 14th century originated from a variant for which marmots in China were the natural reservoir species. [23] [24]
Squirrels are members of the family Sciuridae, a family that includes small or medium-sized rodents. The squirrel family includes tree squirrels, ground squirrels, and flying squirrels. Squirrels are indigenous to the Americas, Eurasia, and Africa, and were introduced by humans to Australia. The earliest known fossilized squirrels date from the Eocene epoch, and among other living rodent families, the squirrels are most closely related to the mountain beaver and dormice.
The groundhog, also known as the woodchuck, is a rodent of the family Sciuridae, belonging to the group of large ground squirrels known as marmots. A lowland creature of North America, it is found through much of the Eastern United States, across Canada and into Alaska. It was first scientifically described by Carl Linnaeus in 1758.
Ground squirrels are rodents of the squirrel family (Sciuridae) that generally live on the ground or in burrows, rather than in trees like the tree squirrels. The term is most often used for the medium-sized ground squirrels, as the larger ones are more commonly known as marmots or prairie dogs, while the smaller and less bushy-tailed ground squirrels tend to be known as chipmunks.
Spermophilus is a genus of ground squirrels in the squirrel family. As traditionally defined the genus was very species-rich, ranging through Europe, Asia and North America, but this arrangement was found to be paraphyletic to the certainly distinct prairie dogs, marmots, and antelope squirrels. As a consequence, all the former Spermophilus species of North America have been moved to other genera, leaving the European and Asian species as true Spermophilus.
The hoary marmot is a species of marmot that inhabits the mountains of northwest North America. Hoary marmots live near the tree line on slopes with grasses and forbs to eat and rocky areas for cover.
The Vancouver Island marmot is a species of marmot endemic to Vancouver Island, in British Columbia. It is one of only five land mammals endemic to Canada.
The alpine marmot is a large ground-dwelling squirrel, from the genus of marmots. It is found in high numbers in mountainous areas of central and southern Europe, at heights between 800 and 3,200 m (2,600–10,500 ft) in the Alps, Carpathians, Tatras and Northern Apennines. In 1948 they were reintroduced with success in the Pyrenees, where the alpine marmot had disappeared at end of the Pleistocene epoch.
The Olympic marmot is a rodent in the squirrel family, Sciuridae. It occurs only in the U.S. state of Washington, at the middle elevations of the Olympic Peninsula. The closest relatives of this species are the hoary marmot and the Vancouver Island marmot. In 2009, it was declared the official endemic mammal of Washington state.
The Himalayan marmot is a marmot species that inhabits alpine grasslands throughout the Himalayas and on the Tibetan Plateau. It is IUCN Red Listed as Least Concern because of its wide range and possibly large population.
The bobak marmot, also known as the steppe marmot, is a species of marmot that inhabits the steppes of Eastern Europe and Central Asia. It is a social animal and inhabits steppe grassland, including cultivated field borders. It hibernates for more than half the year. Litter sizes average about five offspring and it takes three years for the young marmots to reach sexual maturity. Male offspring leave the home colony after their second winter, and about 60% of mature females give birth in any one year.
The mountain ground squirrel is a rodent that is native to southwestern Angola, western Namibia, and western South Africa. It is also known as the Kaoko ground squirrel or the Damara ground squirrel.
The long-tailed marmot or golden marmot is a marmot species in the family Sciuridae. It occurs in mountainous regions in the central parts of Asia where it lives in open or lightly wooded habitats, often among rocks where dwarf junipers grow. It is IUCN Red Listed as Least Concern. As suggested by its name, it is a relatively long-tailed species of marmot.
The gray marmot, grey marmot, or Altai marmot is a species of rodent in the squirrel family Sciuridae. It is one of the larger marmots in the genus Marmota. It occurs in mountainous grasslands and shrub lands of central Asia, and is one of the 9 Palearctic (Eurasia) species. It is found in Xinjiang Province in China, southeastern Kazakhstan, Kyrgyzstan, Mongolia, and in the Altai and Tien Shan Mountains in southeastern Siberia in Russia. In the Mongolian Altai, its range overlaps with that of the Tarbagan marmot. Gray marmots form social groups, live in burrows, and hibernate.
The Alaska marmot, also known as the Brooks Range marmot or the Brower's marmot, is a species of rodent in the family Sciuridae. Once considered to be the same species as the hoary marmot, it is now known to be unique. Alaska marmots are found in the scree slopes of the Brooks Range, Alaska. Specifically, they prefer to dwell on rocky, mountainous terrain, generally near lakes. They eat vegetation found on mountainsides, such as grasses, seeds, and lichen. Their relatively thick bodies are covered in dense, grey fur. They live in large colonies that consist of multiple families. During the winter, they hibernate for long periods of time in burrows. While not well researched, they are not believed to be particularly threatened, by human activity or otherwise. The Alaskan government has designated February 2 as "Marmot Day," a holiday intended to recognize the prevalence of marmots in the state, similar to the more widely celebrated American holiday of Groundhog Day.
The black-capped marmot is a species of rodent in the family Sciuridae. It is endemic to the Russian Far East, but its range is discontinuous and divided into three main parts, each with its own subspecies. The black-capped marmot lives in arctic tundra and alpine habitats from near sea-level to an altitude of 2,000 m (6,600 ft). Depending on exact subpopulation, they hibernate for 6–8 months each year, which is long for a marmot.
The Menzbier's marmot is a species of rodent in the family Sciuridae from Central Asia. Its name commemorates Russian zoologist Mikhail Aleksandrovich Menzbier.
The tarbagan marmot is a species of rodent in the family Sciuridae. It is found in China, northern and western Mongolia, and Russia. In the Mongolian Altai Mountains, its range overlaps with that of the Gray marmot. The species was classified as endangered by the IUCN in 2008.
Hodgson's giant flying squirrel is a species of rodent in the family Sciuridae. This large flying squirrel lives in Himalayan forests in Asia. Like other flying squirrels, it is nocturnal and able to glide long distances between trees by spreading out its patagium, skin between its limbs.
The Xerinae comprise a subfamily of squirrels, many of which are highly terrestrial. It includes the tribes Marmotini, Xerini, and Protoxerini.
The forest-steppe marmot is a rodent species of the marmot genus found in south-central Russia. It lives in wooded forest steppe at an altitude of 180–450 m (590–1,480 ft) in a relatively small region located directly east of the upper Ob River. It has traditionally been considered a subspecies of the similar, more southerly distributed gray marmot, but was separated mainly due to different diploid numbers.