The Wannagan Creek site is a fossil site found just west of the South Unit of Theodore Roosevelt National Park of North Dakota, US. The site is Paleocene in age, approximately 60 million years old. Paleontologists of the Science Museum of Minnesota have studied the site for nearly thirty years. The site is thought to represent a paleoenviroment of subtropical swampy lowland and forests. Preservation is excellent for both the flora and fauna of the site. Trace fossils of crocodilians and other vertebrates have also been discovered.
The topography of the site is that of a badland, due to the downcutting of the Little Missouri River. The rock units are likely sediments derived from the Laramide orogeny deposited in an ecosystem dominated by rivers, streams, ponds, lakes, and swamps.
The Wannagan Site is found in the upper portion of the Tongue River Formation (formerly Bullion Creek Formation). The rocks of this deposit, reaching a thickness of over 60 m, consist of yellow to tan, poorly lithified claystones, mudstones, and siltstones with a lesser amounts of interbedded fine-grained sandstones and lignite. The site is overlain by the Sentinel Butte Formation. This formation is generally gray to brown in color but is similar in lithology to the Tongue River. The Sentinel Butte is roughly 90 m thick. The contact between the two formations is roughly 6 m above the site. The Sentinel Butte contains an extensive stump-bearing petrified wood bed above the contact at the site.
Many mammals are found at Wannagan. All are small compared to modern mammals, the largest being sheep-sized. Preserved mammals include a squirrel-like primitive primate and Ptilodus , an arboreal multituberculate. The largest mammal at the site is the condylarth herbivore Phenacodus . An additional mammal fossil is Protictis , a weasel sized member of Viverridae that may have hunted on the forest floor.
Reptiles are the dominant animal type preserved at Wannagan. The largest animal (~4 meters [13 ft] in length) found at the site is the eusuchian crocodilian Borealosuchus formidabilis (formerly known as Leidyosuchus formidabilis). B. formidabilis is also thought to be the apex predator of the fauna. The second largest reptile is the champsosaur Champsosaurus gigas. C. gigas is unusual among Paleocene reptiles in that it is larger than its known Mesozoic ancestors: 3 meters (10 ft) in length versus 1.5 meters (4.5 ft) for the largest Cretaceous champsosaurs. Reptiles as a whole decreased in size after the Cretaceous–Paleogene extinction event. Other reptiles include soft-shelled turtles, varanid lizards, a small alligatorid called Wannaganosuchus , palaeophid snakes, and the snapping turtle-like Protochelydra zangerli.
Birds from Wannagan include an ibis, an unnamed water bird, and a plover-like shore bird. The two types amphibians found at Wannagan are the giant salamander Piceorpeton willwoodense and frogs. Fish include fossil Esox , freshwater rays, gar, and bowfin. Additionally, the dragonfly Gomphaeschna schrankii and the mayfly Ephemeropteran have been found.
The flora of Wannagan is quite well preserved in some cases, with evidence of insect feeding found on many fossils. Most of the fossils have modern equivalents. The list of trees at Wannagan Creek include palms, ginkgo trees, bald cypress, dogwoods, oaks, elms, fig trees, magnolias, dawn redwoods, hackberries, sassafras, sycamores, cherries, mulberries, and cycads. Additional plants found at the site are grape vines, lotus, bur-reeds, cattails, and water-plantago.
The Chelydridae is a family of turtles that has seven extinct and two extant genera. The extant genera are the snapping turtles, Chelydra and Macrochelys. Both are endemic to the Western Hemisphere. The extinct genera are Acherontemys, Chelydrops, Chelydropsis, Emarginachelys, Macrocephalochelys, Planiplastron, and Protochelydra.
Fossil Butte National Monument is a United States National Monument managed by the National Park Service, located 15 miles (24 km) west of Kemmerer, Wyoming, United States. It centers on an assemblage of Eocene Epoch animal and plant fossils associated with Fossil Lake—the smallest lake of the three great lakes which were then present in what are now Wyoming, Utah, and Colorado. The other two lakes were Lake Gosiute and Lake Uinta. Fossil Butte National Monument was established as a national monument on October 23, 1972.
Deinosuchus is an extinct genus of alligatoroid crocodilian, related to modern alligators and caimans, that lived 82 to 73 million years ago (Ma), during the late Cretaceous period. The name translates as "terrible crocodile" and is derived from the Greek deinos (δεινός), "terrible", and soukhos (σοῦχος), "crocodile". The first remains were discovered in North Carolina in the 1850s; the genus was named and described in 1909. Additional fragments were discovered in the 1940s and were later incorporated into an influential, though inaccurate, skull reconstruction at the American Museum of Natural History. Knowledge of Deinosuchus remains incomplete, but better cranial material found in recent years has expanded scientific understanding of this massive predator.
Champsosaurus is an extinct genus of crocodile-like choristodere reptile, known from the Late Cretaceous and early Paleogene periods of North America and Europe (Campanian–Paleocene). The name Champsosaurus is thought to come from champsai, (χαμψαι) said in an Ancient Greek source to be an Egyptian word for "crocodiles", and sauros, (σαύρος) Greek for "lizard". The morphology of Champsosaurus resembles that of gharials, with a long, elongated snout. It was native to freshwater environments where it likely preyed on fish, similar to living gharials.
Thescelosaurus is an extinct genus of neornithischian dinosaur that lived during the Late Cretaceous period in North America. It was among the last of the non-avian dinosaurs to appear before the entire group went extinct during the Cretaceous–Paleogene extinction event around 66 million years ago. Adult Thescelosaurus would have measured roughly 3–4 metres (10–13 ft) long and probably weighed several hundred kilograms. The genus Thescelosaurus is the type genus and also the largest member of the eponymous Thescelosauridae, which includes similarly-sized bipedal herbivores from the Late Cretaceous of Asia and North America such as Orodromeus, Parksosaurus, and Haya.
The Hell Creek Formation is an intensively studied division of mostly Upper Cretaceous and some lower Paleocene rocks in North America, named for exposures studied along Hell Creek, near Jordan, Montana. The formation stretches over portions of Montana, North Dakota, South Dakota, and Wyoming. In Montana, the Hell Creek Formation overlies the Fox Hills Formation. The site of Pompeys Pillar National Monument is a small isolated section of the Hell Creek Formation. In 1966, the Hell Creek Fossil Area was designated as a National Natural Landmark by the National Park Service.
Cimolestes is a genus of early eutherians with a full complement of teeth adapted for eating insects and other small animals. Paleontologists have disagreed on its relationship to other mammals, in part because quite different animals were assigned to the genus, making Cimolestes a grade taxon of animals with similar features rather than a genus of closely related ones. Fossils have been found in North America, South America, Europe and Africa. Cimolestes first appeared during the Late Cretaceous of North America. According to some paleontologists, Cimolestes died out at the start of the Paleocene, while others report the genus from the early Eocene.
The Fort Union Formation is a geologic unit containing sandstones, shales, and coal beds in Wyoming, Montana, and parts of adjacent states. In the Powder River Basin, it contains important economic deposits of coal, uranium, and coalbed methane.
Borealosuchus is an extinct genus of crocodyliforms that lived from the Late Cretaceous to the Eocene in North America. It was named by Christopher Brochu in 1997 for several species that had been assigned to Leidyosuchus. The species assigned to it are: B. sternbergii, the type species, from the Maastrichtian of Colorado, Montana, North Dakota, South Dakota, and Wyoming; B. acutidentatus, from the Paleocene of Saskatchewan; B. formidabilis, from the Paleocene of North Dakota; B. griffithi, from the Paleocene of Alberta; and B. wilsoni, from the Eocene of Wyoming. B. formidabilis is particularly well-known, represented by the remains of many individuals from the Wannagan Creek site in North Dakota. An indeterminate species is known from the Late Cretaceous Demopolis Chalk in Alabama.
Wannaganosuchus is an extinct genus of small alligatorid crocodilian. It was found in Late Paleocene-age rocks of Billings County, North Dakota, United States.
The Nacimiento Formation is a sedimentary rock formation found in the San Juan Basin of western New Mexico. It has an age of 61 to 65.7 million years, corresponding to the early and middle Paleocene. The formation has yielded an abundance of fossils from shortly after the Cretaceous-Paleogene extinction event that provide clues to the recovery and diversification of mammals following the extinction event.
Orthogenysuchus is an extinct genus of caimanine alligatorids. Fossils have been found from the Wasatch Beds of the Willwood Formation of Wyoming, deposited during the early Eocene. The type species is O. olseni. The holotype, known as AMNH 5178, is the only known specimen belonging to the genus and consists of a skull lacking the lower jaws. The braincase is filled in by the matrix and most of the suture lines between bones are indiscernible, making comparisons with other eusuchian material difficult.
The Ravenscrag Formation is a stratigraphic unit of early Paleocene age in the Western Canada Sedimentary Basin. It was named for the settlement of Ravenscrag, Saskatchewan, and was first described from outcrops at Ravenscrag Butte near the Frenchman River by N.B. Davis in 1918.
Paleontology in North Dakota refers to paleontological research occurring within or conducted by people from the U.S. state of North Dakota. During the early Paleozoic era most of North Dakota was covered by a sea home to brachiopods, corals, and fishes. The sea briefly left during the Silurian, but soon returned, until once more starting to withdraw during the Permian. By the Triassic some areas of the state were still under shallow seawater, but others were dry and hot. During the Jurassic subtropical forests covered the state. North Dakota was always at least partially under seawater during the Cretaceous. On land Sequoia grew. Later in the Cenozoic the local seas dried up and were replaced by subtropical swamps. Climate gradually cooled until the Ice Age, when glaciers entered the area and mammoths and mastodons roamed the local woodlands.
Paleontology in South Dakota refers to paleontological research occurring within or conducted by people from the U.S. state of South Dakota. South Dakota is an excellent source of fossils as finds have been widespread throughout the state. During the early Paleozoic era South Dakota was submerged by a shallow sea that would come to be home to creatures like brachiopods, cephalopods, corals, and ostracoderms. Local sea levels rose and fall during the Carboniferous and the sea left completely during the Permian. During the Triassic, the state became a coastal plain, but by the Jurassic it was under a sea where ammonites lived. Cretaceous South Dakota was also covered by a sea that was home to mosasaurs. The sea remained in place after the start of the Cenozoic before giving way to a terrestrial mammal fauna including the camel Poebrotherium, three-toed horses, rhinoceroses, saber-toothed cat, and titanotheres. During the Ice Age glaciers entered the state, which was home to mammoths and mastodons. Local Native Americans interpreted fossils as the remains of the water monster Unktehi and used bits of Baculites shells in magic rituals to summon buffalo herds. Local fossils came to the attention of formally trained scientists with the Lewis and Clark Expedition. The Cretaceous horned dinosaur Triceratops horridus is the South Dakota state fossil.
Paleontology in Wyoming includes research into the prehistoric life of the U.S. state of Wyoming as well as investigations conducted by Wyomingite researchers and institutions into ancient life occurring elsewhere.
The Tongue River Member is the uppermost geologic member of the Fort Union Formation in Montana, North Dakota, Wyoming. The strata are yellow or light-colored massive sandstones and numerous thick coal beds.
The Golden Valley Formation is a stratigraphic unit of Late Paleocene to Early Eocene age in the Williston Basin of North Dakota. It is present in western North Dakota and was named for the city of Golden Valley by W.E. Benson and W.M. Laird in 1947. It preserves significant assemblages of fossil plants and vertebrates, as well as mollusk and insect fossils.
The 20th century in ichnology refers to advances made between the years 1900 and 1999 in the scientific study of trace fossils, the preserved record of the behavior and physiological processes of ancient life forms, especially fossil footprints. Significant fossil trackway discoveries began almost immediately after the start of the 20th century with the 1900 discovery at Ipolytarnoc, Hungary of a wide variety of bird and mammal footprints left behind during the early Miocene. Not long after, fossil Iguanodon footprints were discovered in Sussex, England, a discovery that probably served as the inspiration for Sir Arthur Conan Doyle's The Lost World.
Coniophis is an extinct genus of snakes from the late Cretaceous period. The type species, Coniophis precedes, was about 7 cm long and had snake-like teeth and body form, with a skull and a largely lizard-like bone structure. It probably ate small vertebrates. The fossil remains of Coniophis were first discovered at the end of the 19th century in the Lance Formation of the US state of Wyoming, and were described in 1892 by Othniel Charles Marsh. For the genus Coniophis, a number of other species have been described. Their affiliation is, however, poorly secured, mostly based on vertebrae descriptions from only a few fossils.